Insecta - Coleoptera - Carabidae
Alternative spelling: Chlaenides
Full reference: G. A. Brullé. 1834. Coléoptères. I. Histoire Naturelle des Insectes, Comprenant Leur Classification, Leurs Moeurs, et la Description des Espèces 1-479
Parent taxon: Harpalinae according to R. P. Webster 2016
See also Bouchard et al. 2011 and LeConte 1861
Sister taxa: Abacetini, Aethocarabus, Dryptini, Galeritini, Harpalini, Helluonini, Lebiini, Licinini, Odacanthini, Panagaeini, Pseudomorphini, Pterostichini, Sinis, Sphodrini, Zabrini, Zuphiini, Oodini
Subtaxa: Chlaeniina
Ecology:
Distribution:
• Quaternary of Greenland (1 collection), Japan (5), the Russian Federation (1), Tanzania (1), the United Kingdom (2), United States (3: Illinois, Pennsylvania, Washington)
• Miocene of China (3)
• Oligocene of France (1)
Total: 17 collections including 20 occurrences