Gastropoda - Stylommatophora - Vertiginidae
Parent taxon: Pupilloidea according to P. Bouchet et al. 2017
See also Harzhauser et al. 2015 and Salvador 2015
Sister taxa: Achatinellidae, Cochlicopidae, Draparnaudiidae, Enidae, Partulidae
Subtaxa: Negulus Nesopupinae Vertigininae
Ecology: epifaunal
Distribution:
• Quaternary of Canada (1: Alberta collection), France (2), Greece (1), Italy (3), Portugal (1), United States (8: Arizona, Florida, Kansas, Michigan, Nevada, Texas, Virginia)
• Pliocene to Pleistocene of Egypt (1), Spain (6)
• Pliocene of Spain (3), Turkey (1)
• Miocene of Austria (6), Bosnia and Herzegovina (1), Germany (6), Hungary (1), Italy (5), Poland (4), Spain (2), Turkey (2)
• Oligocene of Belgium (1)
• Eocene of United States (3: Wyoming)
Total: 58 collections including 91 occurrences