Full reference: B. E. Boudinot, E. V. Yan, J. Prokop, X. Z. Luo, and R. G. Beutel. 2022. Permian parallelisms: Reanalysis of †Tshekardocoleidae sheds light on the earliest evolution of the Coleoptera. Systematic Entomology 48:69-96
Parent taxon: Coleoptera according to B. E. Boudinot et al. 2022
Sister taxa: Acmaeodera perreaui, Adelidium, Adikia, Adynasia, Allognosis, Allopliosilpha, Alphacoleoptera, Anancaeon, Anapiptus, Ancylocheira liasina, Anepismus, Angelinella, Anhuistoma, Anhydrophilus, Anomerus, Anypostatus, Apheloodes, Apicasia, Apioderes, Apiopyrenides, Apistotes, Apsychus, Aptilotitus, Archosyne, Auchenophorites, Babuskaya, Bareus, Bathygerus, Bellingera, Bellingeropsis, Biadelater, Blaptoides, Bothroptera, Brachylaimon, Brachytrachelites, Brevispinnatus, Brochocoleus indibili, Brodiola, Bucklandula, Buprestium, Campopsis, Carabicina, Carabiques, Carabocera, Cardioides, Cardiosyne, Cebrionates, Cerylonopsis, Chimaerocoleites, Chlaeniopsis, Chrysomelidium, Chrysomelinae, Chrysomelophana, Cicindeletae, Cistela, Cistelites, Clathropenna, Clerii, Clinomerus, Coccinellophana, Coleopterites, Coleopteron, Coleopteron depressus, Coleopteron interpunctatus, Coleopteron laevigatus, Coleopteron scabriusculus, Colymbetopsis, Coprophagi, Cretaceites, Cretofalselaterium, Cretogermen, Cretohelophorus, Criocerina pristina, Critoderma, Cryptauchenia, Curculidium, Curculionites liasinus, Curculiopsidae, Curculium, Cydnopsis nigromembranacea, Cyphospheron, Darbiopelta, Diachoristes, Diaperialae, Diaperidium, Diatarastus, Diatrypamene, Dicyphelus, Diphymation, Diplocelides, Diplothece, Doggeria, Dysarestus, Ecthlimma, Elateridopsidae, Elateridopsis, Elaterina, Elytrulum, Enamma, Entomocantharus, Entomophagi, Eocoleopteron, Eodromeus viriosus, Episcepes, Epomenus, Erotilenae, Erotylites, Etheridgea, Euchroma arvernica, Eucoleoptera, Eurynotellus, Eurysphinctus, Eusarcantarus, Explansimarginis, Fengningia, Flichea, Fuscicupes, Gastrodelus, Gastroratus, Gemelina, Glaphoptera, Glaphyropterites, Glaphyropterula, Grahamelytron, Grasselites, Gurvanocoleus, Gyrinites, Gyrinulopsis, Halticophana, Harpalidium, Heeriaopsis, Helophoropsis, Helophorus exilis, Helophorus magnus, Helopidium, Helopii, Helopium, Helops atticus, Hubeicoleus, Hydrobius godeti, Hydrobius obsoletus, Hydrocanthares, Hydroicetes, Hydrophilites, Hydrophilus stenopterus, Hydroporopsis, Hydrous ebeninus, Hylesinus lineatus, Hyperomima, Ipidae, Isomira aurora, Kamaroma, Katapiptus, Keleusticus, Kibdelia, Kryptophagus, Laimocenos, Lamiites, Lamiophanes, Laodiscis, Latridiites, Leioodes, Leptilostis, Leptomites, Leptopodocoleidae, Leptosolenophorus, Longimaxilla, Loxocamarotus, Macrogastri, Macrotrachelites, Malacodermi, Malacodermidae, Masselytron, Megacentrus, Megachorites, Megopis lineolatus, Melanophilites, Melanophilopsis, Menephiloides, Mesoncus, Mesorhynchophora, Mesotricupes, Mesotylites, Metanastes, Metaxycoleoptera, Micrelaterium, Microcoleopteron, Microentomus, Mimelater, Nannoodes, Nebrioides, Necrophagi, Necticus minutus, Nitidulariae, Nubili, Omogongylus, Onkedodimus, Onthophagus luteus, Ooidellus, Ooperiglyptus, Ooperioristus, Oropseini, Oxycephalites, Oxytoroptera, Pachycoleon, Pachypleurites, Paklapwania, Palaeotrachys, Palaeotylinae, Pallax, Palpatores, Pantodapus, Parabuprestium, Paracurculium, Paragrilium, Paragyrinus, Parasilphites, Parnidium, Parnosoma, Pedinosoma, Peltocoleops, Periboloptera, Peridosoma, Petalocupes, Petrorophus, Phaulogyrinus, Pholipheron, Phytoplesion, Pimeliodes, Plastobuprestites, Plastonebria, Platyprosopi, Pleuralocista, Polypamon, Praedodromeus, Prionophana, Prochrysomela, Proheuristes, Prophasis, Prosthecogmus, Prosthenostictus, Protocoleoptera, Protocuneus, Prototoma, Pseudobuprestites, Pseudocarabites, Pseudocurculionites, Pseudocymindis, Pseudocyphon, Pseudoelateropsis, Pseudohydrophilites, Pseudoprionites, Pseudotelephorus, Pseudotenebrio, Pseudus, Rhomaleus, Rhynchophori, Saperda valdensis, Saprophagi, Scalopoides, Semiglobus, Sideriosemion, Silphidium, Sitonites, Smodicoptera, Sojanocoleidae, Sphaericites, Staphyliniae, Stenelytron, Sternoxi, Stictulus, Strongylites, Strullendorfia, Telephorium, Tenebrionites, Tetragonides, Thurmannia, Toksunius, Tolype, Triadocoleopteron, Trialarva, Trichelepturgetes, Tridigitati, Tripsalis, Trixagites, Trochiscites, Trochmalus, Trogossitarii, Trypocoleus, Tshekardocoleidae, Tychon, Ulomites, Velenovskya, Versicolores, Wollastonia, Xenogyrinus, Xinbinia, Xylophagi, Xylotupia, Yanqingia, Zacoleoptera, Zetemenos, Tachypus
Subtaxa: Metacoleoptera Permocupedidae
Ecology: