Parent taxon: Filicopsida according to C. A. Lapasha and C. N. Miller 1985
Sister taxa: Achradosporis, Apiculatasporites, Chelepteris, Choffatia, Cingulatisporites, Clavatisporites, Clavatriletes, Cyathidites, Deltoidospora, Echinatisporis, Foveasporis, Granulatisporites, Grapnelispora, Interlobulites, Ischyosporites, Kirchheimerisporites, Leiotriletes, Lepidodendrites, Marattisporites, Matonisporites, Monolites, Osmundacidites, Pteridacidites, Scabramonoletes, Todisporites, Triletes, Triloboxylon, Waltzispora, Kyrtomisporis, Lophosoriaceae, Lygodiidites, Striatella, Capsispora, Ophioglossidae, Reticuloidosporites, Psilotidae, Polypodiaceoisporites, Lygodioisporites, Vittariaceae, Filicales, Magnastriatites, Cheiropleuriaceae, Lametatriletes, Crassoretitriletes, Hymenophyllopsidaceae, Monachosoraceae
Subtaxa: Arcellites
Distribution:
• Cretaceous of Argentina (12 collections), Australia (25), Canada (96: Alberta, British Columbia, Northwest Territories, Nunavut), China (9), Germany (1), Greenland (1), Poland (3), the United Kingdom (2), United States (55: Colorado, Iowa, Kansas, Minnesota, Montana, Nebraska, New Jersey, Oklahoma, Virginia, Wyoming)
Total: 204 collections including 239 occurrences