Mammalia - Carnivora - Ursidae
Parent taxon: Ursinae according to R. M. Hunt 1998
See also Jin et al. 2007
Sister taxa: Arctotheriini, Aurorarctos, Ballusia, Melursus, Protarctos, Tremarctini, Ursavini
Subtaxa: Ursus
Ecology: ground dwelling herbivore-carnivore
Distribution:
• Quaternary of Austria (1 collection), Belgium (2), Canada (55: British Columbia, Manitoba, Newfoundland, Northwest Territories, Nunavut, Ontario, Quebec, Yukon), China (20), Croatia (2), Czechia (3), Denmark (1), France (24), Georgia (2), Germany (20), Greece (16), Greenland (33), Hungary (6), India (2), Indonesia (1), Israel (3), Italy (36), Japan (4), Kazakhstan (1), Laos (1), Libya (1), Malaysia (2), Mexico (2), Monaco (3), Mongolia (1), Morocco (4), the Netherlands (4), North Atlantic (1), North Pacific (1), Norway (8), Portugal (2), Romania (5), the Russian Federation (10), Spain (23), Svalbard and Jan Mayen (1), Sweden (10), Taiwan (1), Thailand (2), Türkiye (3), the United Kingdom (13), United States (106: Alabama, Alaska, Arkansas, California, Colorado, Florida, Georgia, Idaho, Illinois, Indiana, Iowa, Kentucky, Louisiana, Maine, Maryland, Massachusetts, Michigan, Mississippi, Montana, Nevada, New Mexico, New York, Ohio, Pennsylvania, Tennessee, Texas, Virginia, Washington, West Virginia, Wyoming), Vietnam (1)
• MN 16 of France (2), Italy (1)
• Villafranchian of Greece (2), Hungary (4), Spain (1)
• Blancan of United States (2: California, Texas)
• Pliocene to Pleistocene of China (1), France (2), Greece (2), Hungary (1), Italy (2), Türkiye (1)
• Pliocene of Bulgaria (1), China (1), France (1), Georgia (2), Greece (4), Hungary (2), Italy (1), Moldova (3), Morocco (1), Poland (1), Romania (2), the Russian Federation (3), Spain (1), Tajikistan (4), Ukraine (2), the United Kingdom (2)
• Neogene of Italy (1)
• Miocene of Italy (1)
Total: 491 collections including 526 occurrences