Full reference: A. Lameere. 1900. Notes pour la classification des Coléoptères. Annales de la Société Entomologique de Belgique 44:355-377
Parent taxon: Adephaga according to R. A. Crowson 1981
Sister taxa: Amarodes, Amphizoidae, Amphoxyne, Carabidae, Chimaerocoleus, Cicindelidae, Colymbotethidae, Coptoclavidae, Curculionites, Dytiscidae, Dytiscoidea, Gyrinidae, Haliplidae, Hygrobiidae, Liadytidae, Longurius, Memptus, Noteridae, Oryctites, Parahygrobiidae, Pseudomorphinae, Trachypachidae, Triadogyrus, Triaplidae, Tritarsusidae
Subtaxa: Caraboidea
Ecology:
Distribution:
• Cretaceous of China (16 collections)
• Jurassic of China (1), Germany (5), Kazakhstan (1), Mongolia (2), Poland (1), the Russian Federation (1)
• Triassic of Germany (1)
Total: 28 collections including 42 occurrences