Family Psephenidae Lacordaire 1854 (water penny beetle)

Insecta - Coleoptera - Psephenidae

Parent taxon: Dryopoidea according to C. Y. Cai et al. 2022

See also Bouchard et al. 2011, Crowson 1981, Lawrence and Newton 1995 and Webster 2016

Sister taxa: Actinocerites, Brachypodites, Callirhipidae, Chelonariidae, Cneoglossidae, Dryopidae, Elmidae, Eulichadidae, Expansifemoria, Heteroceridae, Jiuquanocoleus, Limnichidae, Lutrochidae, Mastigocoleidae, Ptilodactylidae

Subtaxa: Eubrianacinae Eubriinae Protacnaeus Psephenella Psepheninae Psephenoidinae Pseudomataeopsephus

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Ecology:

Distribution:

• Quaternary of Japan (2 collections)

• Miocene of Japan (1)

• Oligocene of Brazil (2)

• Eocene of France (1), Germany (3), United States (2: Colorado)

• Cretaceous of Myanmar (4)

Total: 15 collections including 19 occurrences

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