PBDB Taxon

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Leptauchenia lullianus

Mammalia - Merycoidodontidae

Taxonomy

Cyclopidius lullianus was named by Thorpe (1921). Its type locality is Big Muddy Creek, which is in an Arikareean terrestrial horizon in Wyoming.

It was recombined as Pseudocyclopidius lullianus by Schultz and Falkenbach (1968); it was synonymized subjectively with Leptauchenia major by CoBabe (1996), Lander (1998); it was recombined as Leptauchenia lullianus by Stevens and Stevens (2007), Prothero and Sanchez (2008).

Synonyms

Pseudocyclopidius lullianus expiratus was named by Schultz and Falkenbach (1968). Its type specimen is F:AM 45598, a partial skull, and it is a 3D body fossil.

It was synonymized subjectively with Leptauchenia lullianus by Prothero and Sanchez (2008).
Cyclopidius quadratus was named by Koerner (1940). Its type specimen is YPM 13960, a partial skull, and it is a 3D body fossil. Its type locality is West of White Sulphur Springs, which is in a Geringian terrestrial horizon in the Deep River Formation of Montana.

It was recombined as Pseudocyclopidius quadratus by Schultz and Falkenbach (1968); it was synonymized subjectively with Leptauchenia major by CoBabe (1996); it was synonymized subjectively with Leptauchenia loganensis by Lander (1998); it was synonymized subjectively with Leptauchenia lullianus by Prothero and Sanchez (2008).

Synonymy list

Year	Name and author
1921	Cyclopidius lullianus Thorpe
1937	Cyclopidius lullianus Thorpe p. 246 figs. Figs. 173-177
1940	Cyclopidius quadratus Koerner
1968	Pseudocyclopidius lullianus Schultz and Falkenbach
1968	Pseudocyclopidius quadratus Schultz and Falkenbach
1968	Pseudocyclopidius lullianus expiratus Schultz and Falkenbach p. 342 figs. Figures 36, 37, 39, 40, 53
2007	Leptauchenia lullianus Stevens and Stevens p. 158
2008	Leptauchenia lullianus Prothero and Sanchez

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()

Rank	Name	Author
—	Mammaliamorpha
—	Mammaliaformes
class	Mammalia
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Boreoeutheria
—	Laurasiatheria
—	Scrotifera
—	Euungulata
—	Artiodactylamorpha
—	Artiodactyla	()
suborder	Tylopoda	()
family	Merycoidodontidae
subfamily	Leptaucheniinae	Schultz and Falkenbach 1940
genus	Leptauchenia
species	lullianus	(Thorpe 1921)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Leptauchenia lullianus Thorpe 1921

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Invalid names: Cyclopidius quadratus Koerner 1940 [synonym], Pseudocyclopidius lullianus expiratus Schultz and Falkenbach 1968 [synonym]

Diagnosis

Reference	Diagnosis
D. R. Prothero and F. Sanchez 2008	Large leptaucheniines (P2-M3 length = 62-71 mm, M1-3 lengths = 42-48 mm, M1-2 lengths = 27-35 mm, and skull lengths = 125-185 mm) with very robust zygomatic arches and larger orbits; snout relatively flattened and shovel-like; malar relatively deep; teeth very hypsodont.