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Cornwallius sookensis

Mammalia - Perissodactyla - Desmostylidae

Taxonomy
Desmostylus sookensis was named by Cornwall (1922). Its type specimen is BCPM 486, a tooth, and it is a 3D body fossil. Its type locality is Kirby Creek, east, which is in a Chattian marine sandstone in the Sooke Formation of Canada.

It was recombined as Cornwallius sookensis by Hay (1923), Hay (1930), VanderHoof (1937), VanderHoof (1942), Reinhart (1959), Shikama (1966), Domning (1996), Domning (2008), Beatty (2009), Matsui and Tsuihiji (2019), Beatty (2023).

Synonymy list
YearName and author
1922Desmostylus sookensis Cornwall p. 122
1923Desmostylus sookensis Clark and Arnold p. 177
1923Cornwallius sookensis Hay p. 106
1930Cornwallius sookensis Hay p. 654
1937Cornwallius sookensis VanderHoof p. 191
1942Cornwallius sookensis VanderHoof p. 301
1959Cornwallius sookensis Reinhart p. 66
1966Cornwallius sookensis Shikama p. 156
1996Cornwallius sookensis Domning p. 394
2008Cornwallius sookensis Domning p. 642
2009Cornwallius sookensis Beatty p. 896
2019Cornwallius sookensis Matsui and Tsuihiji
2023Cornwallius sookensis Beatty

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
—Therapsida()
infraorderCynodontia()
—Mammaliamorpha
RankNameAuthor
—Mammaliaformes
classMammalia
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—Laurasiatheria
—Scrotifera
—Euungulata
—Panperissodactyla
superorderPerissodactylamorpha
orderPerissodactyla()
orderDesmostylia
orderDesmostyloideaAbel 1933
familyDesmostylidae
genusCornwallius
speciessookensis()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Cornwallius sookensis Cornwall 1922
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Diagnosis
ReferenceDiagnosis
B. L. Beatty 2009Apomorphies of taxon: dental formula is 2.1.2?.3/1.1.1?.3;.
Differs from Behemotops in: molars with longitudinal rows of
columnar cusps that are closely appressed to each other; molars
lacking labial or lingual cingula; molars, specifically M3 and m3,
have styles/stylids in between the main cusps/cuspids, not large
enough to qualify as extra cusps themselves, and these styles/
stylids vary in size between left and right teeth; postorbital process
of the jugal; coronoid process of mandible is more strongly
posterior inclined. Differs from Paleoparadoxia in: molars lacking
labial or lingual cingula; molars with thicker enamel; postorbital
process of the jugal; juvenile mandibles have a posterior
thickening of the symphysis; coronoid process of mandible is
more strongly posterior inclined. Differs from Desmostylus in:molars brachydont; postorbital process does not contact the
zygomatic process of the squamosal since the zygomatic process
of the squamosal is not dorsoventrally expanded.