PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Hustedia pugilla

Rhynchonellata - Athyridida - Neoretziidae

Taxonomy

Hustedia pugilla was named by Cooper and Grant (1976) [Includes four subspecies: hebetata, nasiterna, pluscula, pugilla. Type specimen is holotype for H. pugilla pugilla.]. Its type specimen is USNM 154473b, a shell (Articulated shell), and it is not a trace fossil.

Subspecies

H. hebetata, H. nasiterna, H. pluscula, H. pugilla

Sister species lacking formal opinion data

H. gibbosa, H. grandicosta, H. grandicosta, H. indica, H. indica, H. indica, H. indica, H. marmoni, H. miseri, H. mormonii, H. pseudocardium, H. pygmaea, H. radialis, H. radialis, H. rotundi, H. ulothrix, H. miseri

Synonymy list

Year	Name and author
1976	Hustedia pugilla Cooper and Grant p. 2790 figs. Pl 738, figs 24-68; pl 739, figs 1-34; pl 740, figs 39-42

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
—	Lophophorata
—	Panbrachiopoda	Carlson and Cohen 2020
phylum	Brachiopoda	Cuvier 1805

Rank	Name	Author
subphylum	Rhynchonelliformea	Williams et al. 1996
class	Rhynchonellata	Williams et al. 1996
order	Athyridida	Boucot et al. 1964
suborder	Retziidina	Boucot et al. 1964
superfamily	Retzioidea	Waagen 1883
family	Neoretziidae	Dagis 1972
subfamily	Hustediinae	Grunt 1986
genus	Hustedia	Hall and Clarke 1893
species	pugilla

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Hustedia pugilla Cooper and Grant 1976

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†Hustedia pugilla nasiterna Cooper and Grant 1976

†Hustedia pugilla pluscula Cooper and Grant 1976

†Hustedia pugilla pugilla Cooper and Grant 1976

†Hustedia pugilla hebetata Cooper and Grant 1976

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Diagnosis

No diagnoses are available

Measurements

part	N	mean	minimum	maximum	median	s.d.
shell height	221	5.38	1.10	16.4	6.10	3.71

Composition:

low Mg calcitep

Environment:

marineuc

Locomotion:

stationarysubo

Attached:

yessubo

Life habit:

low-level epifaunalsubo

Diet:

suspension feedersubo

Vision:

blindc

Created:

2018-11-10 18:06:56

Modified:

2018-11-10 18:06:56

Source: subo = suborder, c = class, p = phylum, uc = unranked clade

References: Nesnidal et al. 2013, Aberhan et al. 2004

Age range: base of the Roadian to the top of the Wordian or 274.40000 to 264.28000 Ma

Collections (34 total)

Time interval	Ma	Country or state	Original ID and collection number
Roadian	274.4 - 266.9	USA (Texas)	Hustedia meekana (108862) Hustedia pugilla (43519 43549 43551 43553 81645 81650 101783 102605 102611 102614 105592 105726 105741 105742 105850 105887 105933 106123) Hustedia pugilla, Hustedia pugilla (43548)
Roadian - Wordian	274.4 - 264.28	USA (Texas)	Hustedia pugilla (102630 106049 106053)
Wordian	266.9 - 264.28	USA (Texas)	Hustedia pugilla (43526 43550 43557 105683 105764 105821 105934 105977 106157 106290) Hustedia pugilla, Hustedia pugilla, Hustedia pugilla (43552)