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Agujaceratops mariscalensis

Reptilia - Ceratopsidae

Taxonomy
Chasmosaurus mariscalensis was named by Lehman (1989). Its type specimen is UTEP P.37.3.086, a partial skull (braincase, left supraorbital horncore,left maxilla, and right dentary), and it is a 3D body fossil. Its type locality is WPA Quarry L3 (TMM), which is in a Judithian interdistributary bay claystone in the Aguja Formation of Texas. It is the type species of Agujaceratops.

It was recombined as Agujaceratops mariscalensis by Lucas et al. (2006), Longrich (2010), Scannella and Horner (2011), Maidment and Barrett (2011), Farke (2011), Loewen et al. (2013), Longrich et al. (2013), Longrich (2013), Holmes and Ryan (2013), Currie and Koppelhus (2014), Rivera-Sylva and Carpenter (2014), Longrich (2014), Konishi (2015), Campbell et al. (2016), Lehman et al. (2017), Tweet and Santucci (2018).

Synonymy list
YearName and author
1989Chasmosaurus mariscalensis Lehman p. 139
1990Chasmosaurus mariscalensis Dodson and Currie p. 612
1990Chasmosaurus mariscalensis Lehman p. 215
1993Chasmosaurus mariscalensis Forster and Sereno p. 14
1996Chasmosaurus mariscalensis Lehman p. 501
2001Chasmosaurus mariscalensis Eberth et al. p. 55
2004Chasmosaurus mariscalensis Dodson et al. p. 496
2005Chasmosaurus mariscalensis Diem and Archibald p. 251
2006Chasmosaurus mariscalensis Benammi and Montellano-Ballesteros p. 94
2006Agujaceratops mariscalensis Lucas et al. p. 370
2007Chasmosaurus mariscalensis Lehman p. 259
2007Chasmosaurus mariscalensis Ryan p. 390
2008Chasmosaurus mariscalensis Hunt and Lehman p. 1129
2008Chasmosaurus mariscalensis Williamson and Weil p. 1220
2009Chasmosaurus mariscalensis Erickson et al. p. 1515
2010Agujaceratops mariscalensis Longrich p. 682
2011Agujaceratops mariscalensis Farke p. 7
2011Agujaceratops mariscalensis Maidment and Barrett p. 2
2011Chasmosaurus mariscalensis Porras-Múzquiz and Lehman p. 264
2011Agujaceratops mariscalensis Scannella and Horner p. 5
2013Agujaceratops mariscalensis Holmes and Ryan p. 18
2013Agujaceratops mariscalensis Loewen et al. p. 501
2013Agujaceratops mariscalensis Longrich p. 56
2013Agujaceratops mariscalensis Longrich et al. p. 24
2014Agujaceratops mariscalensis Currie and Koppelhus p. 1035
2014Agujaceratops mariscalensis Longrich p. 292
2014Agujaceratops mariscalensis Rivera-Sylva and Carpenter p. 171
2015Agujaceratops mariscalensis Konishi p. 617
2016Agujaceratops mariscalensis Campbell et al. p. 3
2017Agujaceratops mariscalensis Lehman et al. p. 670
2018Agujaceratops mariscalensis Tweet and Santucci p. 717

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
classReptilia
subclassEureptilia()
—Romeriida
RankNameAuthor
—Diapsida()
—Archosauromorpha(Huene 1946)
—Crocopoda
—ArchosauriformesGauthier 1986
—Eucrocopoda
—Archosauria()
informalAvemetatarsalia
—Ornithodira
—Dinosauromorpha
—Dinosauriformes
—Dinosauria()
—Ornithischia()
—Genasauria
—Cerapoda
—Marginocephalia
—Ceratopsia()
familyCeratopsidae
subfamilyChasmosaurinae
genusAgujaceratops
speciesmariscalensis()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

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Diagnosis
ReferenceDiagnosis
T. M. Lehman 1989Differs from other species of Chasmosaurus in its short, broad squamosal (length/width=1.95, comparedwith 2.60-3.10 in other species) bearing six very large epoccipitals (7-9 in other species) in adults; maxilla without lateral horizontal shelf; premaxilla without posterodorsal extension; supraorbital horncores longer in adults than in other species of Chasmosaurus.
T. M. Lehman 1996Moderately sized chasmosaurine with long, broad. dorsally curved squamosals with convex lateral margins bearing six to ten pronounced marginal undulations with or without attached epoccipitals, a slender medially indented parietal bar with narrow posterolateral rami enclosing very large triangular parietal fenestrae, anteromedian process of the nasals separating the premaxillae, and large posteriorly curved supraborital horncores arising directly over the orbits.The posterior curvature of the supraorbital horncores, dorsal curvature of the squamosals, and slender construction of the parietal distinguish C. mariscalensis from Pentaceratops sternbergii.
T. M. Lehman et al. 2017Agujaceratops distinguished from A. mavericus in having (1) short squamosal with, (2) six episquamosals (or corresponding episquamosal attachment loci on the squamosal), (3) large triangular epi-ossifications, and (4) postorbital horncores with high basal circumference and high taper relative to length.
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yeso
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: gregariouso
Environment: terrestrialsubo
Locomotion: actively mobileo
Life habit: ground dwellingsubo
Diet: herbivoresubo
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2005-09-15 11:08:09
Modified: 2005-09-15 13:08:09
Source: subo = suborder, o = order
References: Marsh 1875, Norman and Weishampel 1991

Age range: Judithian or 83.60000 to 72.20000 Ma

Collections (13 total)


Time interval Ma Country or state Original ID and collection number
Judithian83.6 - 72.2USA (Texas) Chasmosaurus mariscalensis (type locality: 48941 52462 52463 52464)
Late/Upper Campanian83.6 - 72.2USA (Texas) Agujaceratops mariscalensis (236770 236774)
Late/Upper Campanian83.6 - 72.2Mexico (Chihuahua) Chasmosaurus mariscalensis (227230)
Middle Campanian83.6 - 72.2USA (Texas) Agujaceratops mariscalensis (236772 236773 236775)
Middle Campanian - Late/Upper Campanian83.6 - 72.2Mexico (Chihuahua) Chasmosaurus mariscalensis (35353)
Middle Campanian - Late/Upper Campanian83.6 - 72.2USA (Texas) Agujaceratops mariscalensis (236769) Chasmosaurus mariscalensis (22709)