PBDB Taxon

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Romancheinidae

Gymnolaemata - Cheilostomata - Romancheinidae

Taxonomy

Romancheinidae was named by Jullien (1888).

It was assigned to Cheilostomatida by Lee et al. (1997); to Ascophorina by Gordon and Taylor (1999); to Cheilostomata by Bock (2006); and to Lepralielloidea by Schmidt (2003), Berning (2006), Guha and Gopikrishna (2007), Gordon (2012), Gordon and Taylor (2015), Di Martino and Taylor (2015), Brezina et al. (2021).

Synonyms

Escharellidae was named by Levinsen (1909).

It was synonymized subjectively with Schizoporellidae by Bassler (1953); it was synonymized subjectively with Romancheinidae by Gordon (2012).

It was assigned to Ascophora by Brown (1952), Brown (1958), Pouyet (2000); and to Cheilostomata by Bock (2006).
Exochellinae was named by Bassler (1935). It is extant. Its type is Exochella.

It was reranked as the family Exochellidae by Brown (1952), Bassler (1953), Brown (1956), Brown (1958), Pouyet (2000), Bock (2006); it was considered an invalid subgroup of Romancheinidae by Gordon (2012).

It was assigned to Schizoporellidae by Canu and Bassler (1935); to Ascophora by Brown (1952), Bassler (1953), Brown (1956), Brown (1958), Pouyet (2000); and to Cheilostomata by Bock (2006).
Phylactelliporidae was named by Bassler (1953). It is extant. Its type is Phylactellipora.

It was synonymized subjectively with Romancheinidae by Gordon (2012).

It was assigned to Ascophora by Bassler (1953).
Mucronellinae was named by MacGillivray (1887). It is extant.

It was considered an invalid subgroup of Romancheinidae by Gordon (2012).
Peristomellae was named by Canu and Bassler (1917) [Original rank was informal group]. It is extant.

It was considered an invalid subgroup of Romancheinidae by Gordon (2012).
Perigastrellidae was named by Vigneaux (1949). It is extant.

It was synonymized subjectively with Romancheinidae by Gordon (2012).
Hemicycloporinae was named by Vigneaux (1949). It is extant.

It was considered an invalid subgroup of Romancheinidae by Gordon (2012).
Pleurolyrulinae was named by Vigneaux (1949). It is extant.

It was considered an invalid subgroup of Romancheinidae by Gordon (2012).
Peristomellidae was named by Kluge (1962). It is extant.

It was synonymized subjectively with Romancheinidae by Gordon (2012).
Mucronellidae was named by Levinson (1902). Its type is Mucronella.

It was synonymized subjectively with Romancheinidae by Gordon (2012).

It was assigned to Ascophora by Bassler (1953); and to Cheilostomata by Wienrich (1999).

Synonymy list

Year	Name and author
1887	Mucronellinae MacGillivray
1888	Romancheinidae Jullien
1902	Mucronellidae Levinson
1909	Escharellidae Levinsen
1917	Peristomellae Canu and Bassler
1935	Exochellinae Bassler
1935	Exochellinae Canu and Bassler
1949	Perigastrellidae Vigneaux p. 17
1949	Hemicycloporinae Vigneaux p. 83
1949	Pleurolyrulinae Vigneaux p. 105
1952	Escharellidae Brown
1952	Exochellidae Brown
1953	Exochellidae Bassler p. G205
1953	Mucronellidae Bassler p. G207
1953	Phylactelliporidae Bassler p. G217
1956	Exochellidae Brown
1958	Escharellidae Brown
1958	Exochellidae Brown
1962	Peristomellidae Kluge p. 564
1997	Romancheinidae Lee et al.
1999	Romancheinidae Gordon and Taylor
1999	Mucronellidae Wienrich p. 276
2000	Escharellidae Pouyet
2000	Exochellidae Pouyet
2003	Romancheinidae Schmidt
2006	Romancheinidae Berning p. 62
2006	Escharellidae Bock
2006	Exochellidae Bock
2006	Romancheinidae Bock
2007	Romancheinidae Guha and Gopikrishna
2012	Romancheinidae Gordon p. 9
2015	Romancheinidae Di Martino and Taylor
2015	Romancheinidae Gordon and Taylor
2021	Romancheinidae Brezina et al. p. 21

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
—	Lophophorata
phylum	Bryozoa	Ehrenberg 1831

Rank	Name	Author
class	Gymnolaemata	Allman 1856
order	Cheilostomata	Busk 1859
suborder	Flustrina	(Smitt 1868)
infraorder	Umbonulomorpha	Gordon 1989
superfamily	Lepralielloidea	Vigneaux 1949
family	Romancheinidae	Jullien 1888
family	Romancheinidae	Jullien 1888

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Romancheinidae Jullien 1888

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G. Allerescha Gordon 1989

G. Antarcticaetos Hayward and Thorpe 1988

G. Arctonula Gordon and Grischenko 1994

G. †Balantiostoma Marsson 1887

†Balantiostoma elongata Canu 1911

†Balantiostoma spectabilis Canu 1911

G. †Bathosella Canu and Bassler 1917

†Bathosella biavicularia Guha and Gopikrishna 2007

†Bathosella bulbosa Canu and Bassler 1935

†Bathosella laticella Canu and Bassler 1935

G. †Biporochella Guha and Nathan 1996

G. Bostrychopora Hayward and Thorpe 1988

G. †Bulbipora MacGillivray 1895

†Bulbipora areolata MacGillivray 1895

G. Elleschara Gordon 1984

G. Escharella Gordon 1984

Subg. †Escharella (Dialytostoma) Cossmann 1888

†Escharella ashapuraae Guha and Gopikrishna 2007

†Escharella elongata Canu and Bassler 1935

†Escharella fistula Brown 1952

Escharella octodentata Hincks 1880

†Escharella reussiana Busk 1859

†Escharella serrulata Reuss 1848

Escharella spinosissima Hincks 1881

Invalid names: Mucronella [synonym], Perigastrella Canu and Bassler 1917 [synonym]

G. Escharoides Milne Edwards 1836

Escharoides angela Hutton 1873

†Escharoides angela ligulata Brown 1952

Escharoides coccinea Abildgaard 1806

†Escharoides crassa Gordon and Taylor 1999

†Escharoides cuneiformis Guha and Gopikrishna 2007

†Escharoides duplicata Waters 1881

†Escharoides erecta Canu and Bassler 1935

Escharoides excavata MacGillivray 1860

†Escharoides megalota Reuss 1848

Invalid names: Escharoides carboneli David et al. 1972 [synonym], Lepralia fulgurans Manzoni 1869 [synonym]

†Escharoides multispinosa Guha and Gopikrishna 2007

†Escharoides osburni Brown 1958

†Escharoides suggestus Brown 1952

Invalid names: Peristomella [synonym]

G. Exochella Jullien 1888

†Exochella abigailae Gordon and Taylor 2015

Exochella conjuncta Brown 1952

†Exochella gracilis Gordon and Taylor 1999

†Exochella grandis Canu and Bassler 1935

†Exochella jellyae Brown 1952

†Exochella linearis Gordon and Taylor 2015

†Exochella reidae Gordon and Taylor 2015

Exochella tricuspis Hincks 1881

†Exochella woodae Gordon and Taylor 2015

G. Hellerasca Gordon 1989

G. Hemicyclopora Norman 1894

†Hemicyclopora dissidens Gordon and Taylor 2015

†Hemicyclopora ventricosa Gordon and Taylor 2015

G. †Hemiphylactella Vigneaux 1949

G. †Hippolyrula Vigneaux 1949

G. Hippopleurifera Canu and Bassler 1927

†Hippopleurifera australis Gordon and Taylor 1999

†Hippopleurifera benmoussai El Hajjaji 1992

†Hippopleurifera mcbeanensis Cheetham 1962

†Hippopleurifera sedgwicki Milne Edwards 1836

†Hippopleurifera semicristata Reuss 1848

†Hippopleurifera surgens Manzoni 1875

G. †Hoplocheilina Canu 1911

Invalid names: Acanthionella Canu and Bassler 1917 [synonym]

G. Lageneschara Hayward and Thorpe 1988

G. Metrarabdotomorpha d'Hondt 1983

G. †Obliquostoma Thoelen 1968

G. †Ochetosella Canu and Bassler 1917

†Ochetosella jacksonica Canu and Bassler 1917

†Ochetosella parva Cheetham 1962

†Ochetosella robusta Canu and Bassler 1920

G. †Opaeomorpha Zagorsek 2001

G. †Pachykraspedon Koschinsky 1885

G. †Pleuromucrum Vigneaux 1949

G. Pseudosclerodomus d'Hondt and Schopf 1985

G. Psilopsella Canu and Bassler 1927

G. Ragionula Canu and Bassler 1927

G. Romancheina Jullien 1888

G. Temachia Jullien 1882

Invalid names: Escharellidae Levinsen 1909 [synonym], Exochellinae Bassler 1935 [invalid subgroup], Hemicycloporinae Vigneaux 1949 [invalid subgroup], Mucronellidae Levinson 1902 [synonym], Mucronellinae MacGillivray 1887 [invalid subgroup], Perigastrellidae Vigneaux 1949 [synonym], Peristomellae Canu and Bassler 1917 [invalid subgroup], Peristomellidae Kluge 1962 [synonym], Phylactelliporidae Bassler 1953 [synonym], Pleurolyrulinae Vigneaux 1949 [invalid subgroup]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

intermediate Mg calcitef

Composition 2:

aragonitef

Environment:

marineuc

Locomotion:

stationaryo

Attached:

yeso

Life habit:

epifaunalo

Diet:

suspension feedero

Vision:

blindc

Reproduction:

oviparouso

Brooding:

yeso

Comments:

61.3% calcite with 7.5% MgCO3 on average based on 10 specimens f

Created:

2008-02-27 06:39:18

Modified:

2008-02-27 08:39:18

Source: f = family, o = order, c = class, uc = unranked clade

References: Bock 2006, Smith et al. 2006, Nesnidal et al. 2013

Age range

Maximum range based only on fossils: base of the Early/Lower Campanian to the top of the Holocene or 83.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 72.2 Ma

Collections (120 total)

Time interval	Ma	Country or state	Original ID and collection number
Early/Lower Campanian	83.6 - 72.2	USA (Oklahoma)	Escharoides nomas, Escharoides sp. (90502)
Early/Lower Campanian	83.6 - 72.2	USA (Arkansas)	Escharoides erymnos (90501)
Early/Lower Maastrichtian	72.2 - 66.0	USA (Arkansas)	Escharoides danei (90500)
Maastrichtian	72.2 - 66.0	USA (California)	Balantiostoma sp. (87393)
Danian	66.0 - 61.66	Argentina (Río Negro)	Balantiostoma sp., Balantiostoma spectabilis, Balantiostoma elongata (220766)
Danian	66.0 - 61.66	USA (Alabama)	Bathosella undata (83297)
Danian	66.0 - 61.66	Argentina (Neuquén)	Balantiostoma sp. (220769)
Danian	66.0 - 61.66	Sweden	Balantiostoma vallata (69603)
Danian	66.0 - 61.66	Argentina (La Pampa)	Balantiostoma elongata (220764)
Paleocene	66.0 - 56.0	USA (New Jersey)	Bathosella aspera, Acanthionella simplex (84051)
Thanetian - Ypresian	59.24 - 48.07	New Zealand (Chatham Island)	Hippopleurifera australis, Exochella gracilis, Escharoides crassa (7221)
Early/Lower Eocene	56.0 - 48.07	New Zealand (Chatham Island)	Hemicyclopora ventricosa, Exochella abigailae, Exochella reidae, Exochella woodae, Exochella linearis (172944)
Lutetian	48.07 - 41.03	India (Gujarat)	Bathosella biavicularia, Escharoides cuneiformis, Escharoides multispinosa (177176)
Middle Eocene	48.07 - 37.71	India	Bathosella biavicularia, Escharoides cuneiformis, Escharoides multispinosa (178699)
Bartonian	41.03 - 37.71	USA (Alabama)	Acanthionella oecioporosa (88091) Hippopleurifera incondita, Hippopleurifera costulata, Hippopleurifera crassicollis, Perigastrella trapezoidea, Perigastrella ovoidea (7331) Hippopleurifera rotula (7316) Hippopleurifera rotula, Hippopleurifera moodysbranchensis, Perigastrella oscitans (7317)
Bartonian	41.03 - 37.71	United Kingdom (England)	Mucronella angustocecium (42644)
Bartonian	41.03 - 37.71	USA (Florida)	Escharoides erectus, Hippopleurifera costulata, Perigastrella ovoidea (7332) Hippopleurifera crassicollis, Hippopleurifera incondita (7328) Hippopleurifera crassicollis, Hippopleurifera incondita, Perigastrella depressa (7321) Hippopleurifera incondita, Hippopleurifera crassicollis, Perigastrella tubulosa, Perigastrella ovoidea, Ochetosella jacksonica (7330) Hippopleurifera ligulata, Hippopleurifera moodysbranchensis, Hippopleurifera punctata (7325) Hippopleurifera radicata, Hippopleurifera crassicollis, Hippopleurifera incondita (7322 7324)
Late/Upper Eocene	37.71 - 33.9	USA (Georgia)	Hippopleurifera incondita, Perigastrella elegans, Ochetosella jacksonica (160397) Hippopleurifera mcbeanensis, Ochetosella parva (90479) Perigastrella sp., Hippopleurifera mcbeanensis, Ochetosella parva (90480)
Priabonian	37.71 - 33.9	USA (Mississippi)	Mucronella granulosa, Perigastrella oscitans, Perigastrella rectilineata, Peristomella laticella, Ochetosella jacksonica, Acanthionella oecioporosa (84174)
Priabonian	37.71 - 33.9	USA (Florida)	Hippopleurifera punctata (7318)
Priabonian	37.71 - 33.9	New Zealand (North Otago)	Escharella sp., Escharoides sp., Exochella sp. 1, Romancheinidae indet. (172816) Escharella sp., Exochella sp. 1, Exochella sp. 2, Hemicyclopora sp. (172815)
Runangan	36.7 - 34.6	New Zealand	Escharoides suggestus (173838)
Rupelian	33.9 - 27.3	USA (Mississippi)	Perigastrella plana (90565)
Rupelian	33.9 - 27.3	USA (Alabama)	Perigastrella ovoidea (90551) Perigastrella plana (90563) Perigastrella plana, Perigastrella costellifera, Peristomella coccinea (90561 90562) Perigastrella plana, Peristomella erecta (90549)
Oligocene	33.9 - 23.04	New Zealand (North-West Nelson)	Escharoides angela, Escharella fistula (173837)
Oligocene - Miocene	33.9 - 5.333	Australia (Victoria)	Escharella elongata (173521) Escharella elongata, Bathosella bulbosa, Escharoides duplicata, Escharoides erecta, Escharoides osburni (173508) Escharella elongata, Escharoides duplicata, Escharoides osburni (173512) Escharoides duplicata (173527)
Chattian	27.3 - 23.04	Australia (Victoria)	Bathosella bulbosa, Bathosella laticella (174140) Escharoides erecta (174144 174148 174150 174152) Mucronella elongata (174154)
Late/Upper Oligocene	27.3 - 23.04	New Zealand	Exochella tricuspis (173832)
Miocene - Oligocene	23.04 - 23.04	Australia (Victoria)	Bulbipora areolata, Escharella elongata, Escharoides duplicata (173502) Escharella elongata, Escharoides duplicata (173504 173511) Escharella elongata, Escharoides duplicata, Escharoides osburni (173506)
Aquitanian	23.04 - 20.45	Colombia	Escharoides martae, Hippopleurifera sp. 1, Pleuromucrum sp. (211332) Hippopleurifera sp. 1 (211333) Hippopleurifera sp. 1, Hippopleurifera mucronata (211331)
Aquitanian - Burdigalian	23.04 - 15.98	Colombia	Hippopleurifera sp. 2 (211330)
Early/Lower Miocene	23.04 - 15.98	India	Escharella ashapurae (178702)
Burdigalian	20.45 - 15.98	Egypt	Hippolyrula crassilabiata (47007)
Burdigalian	20.45 - 15.98	France	Escharella sp. (65395)
Altonian	18.7 - 15.9	New Zealand	Escharoides angela (173830)
Langhian	15.98 - 13.82	Australia (Victoria)	Exochella grandis (174139)
Middle Miocene	15.98 - 11.63	Australia (Victoria)	Exochella grandis (166365)
Middle Miocene	15.98 - 11.63	Romania	Escharella tenera, Escharoides coccinea (178089)
Middle Miocene	15.98 - 11.63	Czechia	Escharella chiragra, Escharoides mamillata (178887) Escharella sp., Hippopleurifera sedgwicki, Hippopleurifera semicristata (178897) Escharella tenera, Escharoides aliferus, Escharoides cocinea, Hippopleurifera sedgwicki (178895) Escharella tenera, Escharoides cocinea, Hippopleurifera sedgwicki (178890) Escharella tenera, Escharoides cocinea, Perigastrella concinna, Hippopleurifera sedgwicki, Hippopleurifera semicristata (178899) Escharella tenera, Escharoides grotriani, Hippopleurifera sedgwicki (178896) Escharella variolosa, Escharoides aliferus, Escharoides grotriani (178893) Escharoides megalota (178888) Hippopleurifera sedgwicki (178894 178902 178904) Hippopleurifera sedgwicki, Hippopleurifera semicristata (178892 178901) Perigastrella concinna, Hippopleurifera sedgwicki, Hippopleurifera semicristata (178898)
Serravallian	13.82 - 11.63	Hungary	Hippopleurifera hypsostoma (34739)
Waiauan	13.05 - 11.04	New Zealand	Escharella fistula (173831)
Tortonian	11.63 - 7.246	Spain	Escharoides coccinea (177480) Escharoides coccinea, Escharoides fulgurans (177482) Escharoides sp., Hemicyclopora sp. 1, Hemicyclopora sp. 2, Escharoides coccinea, Escharella reussiana, Escharella serrulata, Escharoides megalota, Escharella peachi, Escharella grossa, Hippopleurifera sedgwicki (178465) Hippopleurifera surgens, Escharoides coccinea, Escharella octodentata (177485) Hippopleurifera surgens, Escharoides coccinea, Escharoides fulgurans (177481) Hippopleurifera surgens, Escharoides coccinea, Escharoides fulgurans, Escharella reussiana (177486)
Messinian	7.246 - 5.333	Algeria	Hippopleurifera sulcimargo (70785)
Messinian	7.246 - 5.333	Spain	Hippopleurifera benmoussai (177487)
Messinian	7.246 - 5.333	Morocco	Escharoides coccinea (34723 34724)
Zanclean	5.333 - 3.6	Portugal	Escharella sp., Escharoides sp., Hippopleurifera sp. (152330)
Pliocene	5.333 - 2.58	New Zealand	Exochella jellyae, Exochella conjuncta, Escharoides angela (173739) Exochella jellyae, Exochella conjuncta, Escharoides angela, Escharella fistula (173737) Exochella jellyae, Exochella conjuncta, Escharoides angela, Escharella spinosissima (173740)
Pliocene	5.333 - 2.58	Australia (South Australia)	Escharoides excavata (172911)
Pliocene	5.333 - 2.58	Antarctica	Exochella longirostris, Mucronella praestans (41581) Exochella sp. (41577) Exochella sp., Hemicyclopora polita (41578) Phylactellipora sp. (41579) Psilopsella uniseriata (41575) Romancheina sp. (41576)
Pliocene	5.333 - 2.58	Spain	Hippopleurifera sedgwicki, Hippopleurifera surgens, Escharoides coccinea, Escharoides fulgurans (177490 177491) Hippopleurifera surgens, Escharoides coccinea, Escharoides fulgurans (177489)
Early/Lower Pleistocene	2.58 - 0.774	Antarctica	Lageneschara lyrulata (48425 48427 48432)
Early/Lower Pleistocene	2.58 - 0.774	Italy	Hemicyclopora multispinata, Neolagenipora collaris (146291)
Pleistocene	2.58 - 0.0117	New Zealand	Escharoides excavata (173728)
Calabrian	1.8 - 0.774	Costa Rica (Limon)	Hippopleurifera n. sp. 2 (79845)
Castlecliffian	1.63 - 0.34	New Zealand	Escharoides excavata (173734)
Middle Pleistocene	0.774 - 0.129	Greenland	Escharella connectens, Escharella indivisa, Escharella klugei (75200) Escharella connectens, Escharella indivisa, Escharella klugei, Escharella thompsoni (75197) Escharella connectens, Escharella klugei (75179)
Late/Upper Pleistocene	0.129 - 0.0117	Greenland	Escharella klugei (75148)
Late/Upper Pleistocene	0.129 - 0.0117	USA (California)	Mucronella microstoma (160361)
Holocene	0.0117 - 0.0	USA (Florida)	Escharoides costifer (97607)