PBDB Taxon

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Permophricodothyris indica

Rhynchonellata - Spiriferida - Elythidae

Taxonomy

Reticularia indica was named by Waagen (1883).

It was recombined as Squamularia indica by Grabau (1931), Huang (1933), Shimizu (1961), Feng and Jiang (1978), Liao (1980), Zeng et al. (1995); it was recombined as Squamularia (Neophricodothyris) indica by Reed (1944); it was recombined as Neophricadothyris indica by Ruzhentsev and Sarycheva (1965); it was recombined as Phricodothyris indicus by Iran-Japanese Research Group (1981); it was recombined as Bullarina indica by Liang (1990); it was recombined as Permophricodothyris indica by Shi et al. (2002), Campi et al. (2005).

Synonyms

Anomaloria dorsatusa was named by Liang (1990). It is a 3D body fossil. Its type locality is Lengwu, upper Shiziling Member, Tonglu, which is in a Capitanian coastal marl/limestone in the Lengwu Formation of China.

It was synonymized subjectively with Permophricodothyris indica by Clapham (2015).
Bullarina pseudoasiatica was named by Liang (1990). It is a 3D body fossil. Its type locality is Lengwu, upper Shiziling Member, Tonglu, which is in a Capitanian coastal marl/limestone in the Lengwu Formation of China.

It was recombined as Permophricodothyris pseudoasiatica by Shi et al. (2002); it was synonymized subjectively with Permophricodothyris indica by Clapham (2015).
Astegosia plenussa was named by Liang (1990). It is a 3D body fossil. Its type locality is Lengwu, basal Shiziling Member, Tonglu, which is in a Capitanian coastal marl/limestone in the Lengwu Formation of China.

It was synonymized subjectively with Permophricodothyris indica by Clapham (2015).
Astegosia qianjiangensis was named by Liang (1990). It is a 3D body fossil. Its type locality is Lengwu, basal Shiziling Member, Tonglu, which is in a Capitanian coastal marl/limestone in the Lengwu Formation of China.

It was synonymized subjectively with Permophricodothyris indica by Clapham (2015).
Astegosia radicula was named by Liang (1990). It is a 3D body fossil. Its type locality is Lengwu, basal Shiziling Member, Tonglu, which is in a Capitanian coastal marl/limestone in the Lengwu Formation of China.

It was synonymized subjectively with Permophricodothyris indica by Clapham (2015).
Astegosia stropharostrisa was named by Liang (1990). It is a 3D body fossil. Its type locality is Lengwu, lower Cunhouling Member, Tonglu, which is in a Capitanian coastal siliciclastic in the Lengwu Formation of China.

It was synonymized subjectively with Permophricodothyris indica by Clapham (2015).
Anomaloria puerilis was named by Liang (1990). It is a 3D body fossil. Its type locality is Lengwu, upper Shiziling Member, Tonglu, which is in a Capitanian coastal marl/limestone in the Lengwu Formation of China.

It was synonymized subjectively with Permophricodothyris indica by Clapham (2015).

Synonymy list

Year	Name and author
1883	Reticularia indica Waagen p. 542 figs. Pl 43, fig 6; pl 44, fig 2
1900	Reticularia indica Arthaber p. 270 figs. Pl 20, figs 4a-d, 5a-d
1913	Reticularia indica Mansuy p. 120 fig. 11
1931	Squamularia indica Grabau p. 200 figs. Pl 17, fig 5; pl 18, figs 2-4
1931	Reticularia indica Reed p. 25 figs. Pl 4, figs 6,6a
1933	Squamularia indica Huang p. 38 figs. Pl 5, figs 8-10
1944	Squamularia (Neophricodothyris) indica Reed p. 234
1961	Squamularia indica Shimizu p. 333 figs. Pl 17, figs 4-7,13
1965	Neophricadothyris indica Ruzhentsev and Sarycheva
1978	Squamularia indica Feng and Jiang p. 296 figs. Pl 105, fig 4
1980	Squamularia indica Liao figs. Pl 8, figs 57,58
1981	Phricodothyris indicus Iran-Japanese Research Group
1990	Anomaloria dorsatusa Liang
1990	Bullarina indica Liang p. 295 figs. Pl 66, figs 8-12
1990	Bullarina pseudoasiatica Liang p. 296 figs. Pl 65, figs 1-12
1990	Astegosia plenussa Liang p. 299 figs. Pl 67, figs 1-11
1990	Astegosia qianjiangensis Liang p. 300 figs. Pl 68, figs 11-20
1990	Astegosia radicula Liang p. 301 figs. Pl 68, figs 1-5
1990	Astegosia stropharostrisa Liang p. 302 figs. Pl 51, figs 15,16; pl 69, figs 1-25; pl 70, figs 1-10
1990	Anomaloria dorsatusa Liang p. 306 figs. Pl 64, figs 11-15
1990	Anomaloria puerilis Liang p. 307 figs. Pl 62, figs 11-15; pl 65, figs 13-17
1995	Squamularia indica Zeng et al. figs. Pl 14, fig 7
2002	Permophricodothyris indica Shi et al. p. 376
2002	Permophricodothyris pseudoasiatica Shi et al. p. 376
2004	Anomaloria dorsatusa Chen p. 46 figs. Pl 5, figs 25-33; pl 9, figs 24-27; pl 10, figs 1-9
2005	Permophricodothyris indica Campi et al. p. 144 figs. Pl 7, figs C, D, P-T, V-aa

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
—	Lophophorata
—	Panbrachiopoda	Carlson and Cohen 2020
phylum	Brachiopoda	Cuvier 1805

Rank	Name	Author
subphylum	Rhynchonelliformea	Williams et al. 1996
class	Rhynchonellata	Williams et al. 1996
order	Spiriferida
suborder	Delthyridina	Ivanova 1972
superfamily	Reticularioidea
family	Elythidae	Fredericks 1924
subfamily	Phricodothyridinae	Caster 1939
genus	Permophricodothyris
species	indica	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Permophricodothyris indica Waagen 1883

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Invalid names: Anomaloria dorsatusa Liang 1990 [synonym], Anomaloria puerilis Liang 1990 [synonym], Astegosia plenussa Liang 1990 [synonym], Astegosia qianjiangensis Liang 1990 [synonym], Astegosia radicula Liang 1990 [synonym], Astegosia stropharostrisa Liang 1990 [synonym], Bullarina pseudoasiatica Liang 1990 [synonym]

Diagnosis

No diagnoses are available

Measurements

part	N	mean	minimum	maximum	median	s.d.
shell height	27	18.5	6.80	36.0	17.6	6.42

Composition:

low Mg calcitep

Environment:

marineuc

Locomotion:

stationaryo

Attached:

yesp

Life habit:

epifaunalo

Diet:

suspension feedero

Vision:

blindc

Created:

2004-02-28 14:12:52

Modified:

2004-02-28 16:12:52

Source: o = order, c = class, p = phylum, uc = unranked clade

References: Nesnidal et al. 2013, Aberhan et al. 2004

Age range: base of the Asselian to the top of the Changhsingian or 298.90000 to 251.90200 Ma

Collections (75 total)

Time interval	Ma	Country or state	Original ID and collection number
Asselian	298.9 - 293.52	China (Xinjiang)	Anomaloria dorsatusa (105228)
Sakmarian	293.52 - 290.1	China (Xinjiang)	Anomaloria dorsatusa (35514)
Sakmarian - Artinskian	293.52 - 283.3	Timor-Leste	Squamularia indica (21163)
Guadalupian	274.4 - 259.51	China (Yunnan)	Spirifer indica (139252)
Wordian	266.9 - 264.28	Pakistan	Squamularia indica (125876)
Wordian	266.9 - 264.28	Azerbaijan	Neophricadothyris indica (82257 113610)
Wordian	266.9 - 264.28	China (Nei Mongol)	Squamularia indica (109168 109179 109180 109209 109213 109214 109257)
Wordian	266.9 - 264.28	Iran	Phricodothyris indicus (112909)
Wordian - Capitanian	266.9 - 259.51	Cambodia	Reticularia indica (125049)
Capitanian	264.28 - 259.51	Pakistan	Reticularia indica (119615)
Capitanian	264.28 - 259.51	China (Zhejiang)	Anomaloria dorsatusa, Bullarina indica, Bullarina pseudoasiatica, Astegosia qianjiangensis, Astegosia stropharostrisa, Anomaloria puerilis (111933) Astegosia plenussa, Astegosia qianjiangensis, Astegosia radicula, Astegosia stropharostrisa, Anomaloria puerilis, Astegosia sp. 1 (111934) Astegosia qianjiangensis, Astegosia stropharostrisa (95519) Astegosia qianjiangensis, Astegosia stropharostrisa, Martinia sp. 1 (111932)
Capitanian	264.28 - 259.51	China (Nei Mongol)	Squamularia indica (109256)
Capitanian	264.28 - 259.51	Malaysia	Permophricodothyris indica (105035)
Wuchiapingian	259.51 - 254.14	China (Guangxi)	Squamularia indica (112366)
Wuchiapingian	259.51 - 254.14	Japan	Squamularia indica (117840)
Wuchiapingian	259.51 - 254.14	China (Jiangxi)	Squamularia indica (101631)
Wuchiapingian	259.51 - 254.14	Pakistan	Reticularia indica (89060 119550 119897 119931 120124 124406) Spirifera lineata (117776) Squamularia indica (125896 125898 125902 125903 125908 125918 126281 126293 126299 126343 126453 126459 126994 127007)
Wuchiapingian	259.51 - 254.14	Azerbaijan	Reticularia indica (132466) Spirifer lineatus (232366)
Wuchiapingian	259.51 - 254.14	China (Sichuan)	Permophricodothyris indica (148473)
Wuchiapingian	259.51 - 254.14	China (Guizhou)	Squamularia indica (108192 110973 112000 112001 112002 112004 136249)
Wuchiapingian	259.51 - 254.14	China (Chongqing)	Squamularia indica (111069 156843 156867 156868 156869)
Lopingian	259.51 - 251.902	China (Jiangxi)	Spirifer lineatus (109628)
Lopingian	259.51 - 251.902	Pakistan	Reticularia indica (236245)
Wuchiapingian - Changhsingian	259.51 - 251.902	Pakistan	Spirifera lineata (117777) Squamularia indica (126695 126704 126990)
Changhsingian	254.14 - 251.902	China (Hubei)	Squamularia indica (120809)
Changhsingian	254.14 - 251.902	China (Xizang)	Squamularia indica (187558)
Changhsingian	254.14 - 251.902	Pakistan	Reticularia indica (90251)
Changhsingian	254.14 - 251.902	China (Chongqing)	Squamularia indica (94984 94985 156855 156858)