PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Cardiella

Cephalopoda - Goniatitida - Marathonitidae

Taxonomy

Cardiella was named by Pavlov (1967) [Sepkoski's age data: P Asse P Leon-m Sepkoski's reference number: 732]. It is not extant.

It was assigned to Goniatitida by Sepkoski (2002); to Marathonitinae by Leonova (1989), Bogoslovskaya (1990), Leonova (2002) and Korn (2006); and to Marathonitidae by Furnish et al. (2009) and Zhou (2017).

Species

Aksuites permicus, C. busterensis, C. ganti, C. gracia (syn. C. amygdala), C. kussica, C. martodjojoi, C. multicostata, C. pamirica, C. parva, C. romanowskyi, C. ruzhencevi, C. shyndensis, C. suevi, C. sulcata, C. vetusta

Synonyms

Aksuites was named by Pavlov (1967).

It was synonymized subjectively with Cardiella by Furnish et al. (2009).

It was assigned to Marathonitinae by Leonova (1989), Bogoslovskaya (1990), Leonova (2002) and Korn (2006).

Synonymy list

Year	Name and author
1967	Aksuites Pavlov
1967	Cardiella Pavlov
1989	Aksuites Leonova p. 146
1989	Cardiella Leonova p. 149
1990	Aksuites Bogoslovskaya p. 72
1990	Cardiella Bogoslovskaya p. 72
2002	Aksuites Leonova
2002	Cardiella Leonova
2002	Cardiella Sepkoski
2006	Aksuites Korn
2006	Cardiella Korn
2009	Cardiella Furnish et al. p. 160
2017	Cardiella Zhou p. 68

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
phylum	Mollusca

Rank	Name	Author
class	Cephalopoda	Cuvier 1797
subclass	Ammonoidea	()
order	Goniatitida	Hyatt 1884
suborder	Goniatitina	Hyatt 1884
superfamily	Marathonitoidea	(Ruzhencev 1938)
family	Marathonitidae	Ruzhencev 1938
genus	Cardiella	Pavlov 1967

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Cardiella Pavlov 1967

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†Cardiella busterensis Toumanskaya 1949

†Cardiella ganti Smith 1903

†Cardiella gracia Pavlov 1967

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Invalid names: Cardiella amygdala Leonova 1981 [synonym]

†Cardiella kussica Toumansky 1931

†Cardiella martodjojoi Glenister and Furnish 1987

†Cardiella multicostata Leonova 1992

†Cardiella pamirica Toumansky 1949

†Cardiella parva Leonova 1981

†Cardiella romanowskyi Karpinsky 1890

†Cardiella ruzhencevi Leonova 1981

†Cardiella shyndensis Leonova 1981

†Cardiella suevi Toumanskaya 1931

†Cardiella sulcata Böse 1919

†Cardiella vetusta Bogoslovskaya 1978

Invalid names: Aksuites Pavlov 1967 [synonym]

Diagnosis

No diagnoses are available

Measurements

part	N	mean	minimum	maximum	median	s.d.
shell height	37	12.8	5.30	29.0	12.5	5.91
shell diameter	37	22.1	9.20	52.0	21.5	10.4
shell inflation	37	0.003	0.001	4.00	0.001	0.716

Composition:

aragoniteo

Environment:

marinec

Locomotion:

fast-movingo

Life habit:

nektonico

Diet:

carnivoreo

Vision:

well-developedc

Created:

2005-01-28 02:36:55

Modified:

2005-10-06 16:36:08

Source: o = order, c = class

References: Kiessling 2003, Kiessling 2004

Age range: base of the Gzhelian to the top of the Wordian or 303.70000 to 264.28000 Ma

Collections (95 total)

Time interval	Ma	Country or state	Original ID and collection number
Gzhelian	303.7 - 298.9	USA (Texas)	Marathonites ganti (43477 43483 43484 43485 128584 128645) Marathonites sulcatus (128582) Marathonites sulcatus, Marathonites ganti (128678) Stacheoceras sulcatus (128585)
Virgilian	303.7 - 298.9	USA (Texas)	Marathonites ganti (217891 217903 217906 217908 217913 217979 217986)
Asselian	298.9 - 295.5	Tajikistan	C. vetusta (102491 102492 102494 102495 102496)
Cisuralian	298.9 - 273.01	Iran	C. sp. (226701)
Sakmarian	293.52 - 290.1	Tajikistan	C. multicostata (130243 130245)
Sakmarian - Artinskian	293.52 - 279.3	Indonesia (Timor)	C. martodjojoi (131192)
Artinskian	290.1 - 279.3	Kazakhstan	Marathonites invariabilis (102148)
Artinskian	290.1 - 279.3	Tajikistan	C. pamirica (130246) C. shyndensis (130247 130284)
Artinskian	290.1 - 279.3	Russian Federation	Marathonites pamiricus (111239)
Kungurian	279.3 - 272.3	Tajikistan	Aksuites permicus (111291 111292 177982) C. amygdala (111259 178037) C. amygdala, C. gracia, C. parva (111258) C. amygdala, C. gracia, C. parva, C. pamirica (111278) C. amygdala, C. gracia, C. parva, C. shyndensis (111260) C. amygdala, C. pamirica (178138) C. amygdala, C. parva (111288 111290) C. gracia (111261 111266 177981 178132) C. gracia, C. pamirica (111277 178047) C. gracia, C. parva (111248 111275 178036) C. gracia, C. parva, C. shyndensis (111283 111285) C. pamirica (111274 111286 178035) C. pamirica, C. shyndensis (111270 178023) C. pamirica, C. shyndensis, Aksuites permicus (111281) C. parva (111247 111249 111250 111251 111253 111254 111255 111256 111257 111262 111263 111264 111265 111267 111268 111269 111273 111276 111287 111289 178033) C. parva, C. pamirica (111272 111279 111378) C. parva, C. shyndensis (111252 111282) C. shyndensis (111271 111280 111284 177983 178028 178133)
Kungurian	279.3 - 272.3	China (Guangxi)	C. gracia (185587)
Roadian	272.3 - 268.8	Ukraine	C. kussica (81423 110830 110835)
Wordian	266.9 - 264.28	Japan	C. sp. (107611)