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Pteraspidiformes

Pteraspidomorpha - Pteraspidiformes

Synonymy list
YearName and author
1940Pteraspidiformes Berg
1996Pteraspidiformes Elliott and Ilyes p. 153
2002Pteraspidiformes Blieck et al. p. 811
2005Pteraspidiformes Dumbrava and Blieck p. 165
2011Pteraspidiformes Long p. 241
2016Pteraspidiformes Nelson et al. p. 29
2017Pteraspidiformes Randle and Sansom p. 596
2019Pteraspidiformes van der Laan p. 17

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
RankNameAuthor
superclassPteraspidomorphi(Goodrich 1909)
classPteraspidomorpha
subclassHeterostraciLankester 1868
orderPteraspidiformesBerg 1940
orderPteraspidiformesBerg 1940

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. †Pteraspidiformes Berg 1940
show all | hide all
Fm. †Anchipteraspididae Elliott 1984
+
G. †Anchipteraspis Elliott 1984
+
†Anchipteraspis crenulata Elliott 1984
G. †Rhachiaspis Elliott 1984
+
†Rhachiaspis pteriga Elliott 1984
†Rhachiaspis pteriga Elliott 1984
Invalid names: Rhachiaspis pteriga Elliott 1984 [valid], Rhachiaspis pteriga Elliott 1984 [valid]
G. †Ulutitaspis Elliott 1984
+
†Ulutitaspis notidana Elliott 1984
G. †Dnestraspis Novitskaya 1983
+
†Dnestraspis firma Novitskaya 1983
Fm. †Protaspidae Bryant 1933
+
G. †Eucyrtaspis White and Moy-Thomas 1940
+
Invalid names: Cyrtaspis Bryant 1932 [replaced]
G. †Protaspis Bryant 1933
+
Subg. †Protaspis (Europrotaspis) White 1961
Subg. †Protaspis (Protaspis) Bryant 1933
+
†Protaspis brevirostris Bryant 1933
†Protaspis bucheri Bryant 1932
†Protaspis cingulus Bryant 1933
†Protaspis constrictus Bryant 1933
Fm. †Protopteraspididae Novitskaya 1983
+
G. †Canadapteraspis Dineley and Loeffler 1976
+
†Canadapteraspis alocostomata Dineley and Loeffler 1976
†Canadapteraspis formosa Thorsteinsson and Elliott 2022
†Canadapteraspis helostomata Dineley and Loeffler 1976
†Canadapteraspis thymostomata Dineley and Loeffler 1976
†Canadapteraspis uniformis Thorsteinsson and Elliott 2022
G. †Escharaspis Elliott 1983
+
†Escharaspis alata Elliott 1983
G. †Gigantaspis Heintz 1962
+
†Gigantaspis isachseni Heintz 1962
†Gigantaspis laticephala PERNEGRE and GOUJET 2007
†Gigantaspis minima PERNEGRE and GOUJET 2007
G. †Loricopteraspis Halstead Tarlo 1961
+
†Loricopteraspis dairydinglensis Ball and Dineley 1961
G. †Miltaspis Blieck 1982
+
†Miltaspis anatirostrata Blieck 1982
G. †Protopteraspis Leriche 1924
+
†Protopteraspis arctica Elliott 1983
†Protopteraspis corniga Elliott 1983
†Protopteraspis gosseleti Leriche 1906
†Protopteraspis micra Ilyes et al. 1995
†Protopteraspis pygmaea Elliott 1983
†Protopteraspis sartokia Elliott 1983
†Protopteraspis siliktokia Elliott 1983
†Protopteraspis vogti Kiaer 1928
Invalid names: Simopteraspis Denison 1970 [synonym]
G. †Stegobranchiaspis Elliott 1983
+
G. †Zascinaspis Stensiu 1958
+
†Zascinaspis heintzi Blieck 1984
†Zascinaspis laticephala Blieck and Goujet 1983
Subor. †Psammosteoidei
+
Fm. †Obrucheviidae Halstead Tarlo 1964
+
G. †Aspidosteus Obruchev 1941
G. †Obruchevia Halstead Tarlo 1964
+
†Obruchevia heckeri Halstead Tarlo 1964
G. †Perscheia Elliott et al. 2004
+
†Perscheia pulla Elliott et al. 2004
Invalid names: Aspidosteidae Berg 1955 [synonym]
Fm. †Pteraspididae Claypole 1885
+
G. †Archaeoteuthis Roemer 1855
G. †Chiastolepis Eichwald 1844
G. †Ecphymaspis Elliott et al. 2020
+
†Ecphymaspis idahoensis Elliott et al. 2020
+
Invalid names: Psephaspis idahoensis Denison 1968 [replaced]
G. †Glossoidaspis Branson and Mehl 1931
+
†Glossoidaspis giganteus Branson and Mehl 1931
G. †Holaspis Lankester 1873
+
†Kallostrakon alleni Halstead Tarlo 1964
†Kallostrakon grindrodi Halstead Tarlo 1964
†Kallostrakon macanuffi Halstead Tarlo 1964
†Kallostrakon podura Lankester 1870
G. †Mitraspis Elliott et al. 2015
+
†Mitraspis cracens Elliott et al. 2015
G. †Mylopteraspis Stensiu 1958
G. †Palaeoteuthis Roemer 1855
+
†Palaeoteuthis marginalis Eichwald 1871
G. †Rhachiaspis Elliott 1984
+
†Rhachiaspis pteriga Elliott 1984
†Rhachiaspis pteriga Elliott 1984
Invalid names: Rhachiaspis pteriga Elliott 1984 [valid], Rhachiaspis pteriga Elliott 1984 [valid]
G. †Scaphaspis Lankester 1865
+
†Scaphaspis nastorthi Lankester 1884
G. †Scutellaspis Elliott et al. 2020
+
†Scutellaspis wilsoni Elliott et al. 2020
G. †Sphagodus Agassiz 1839
Invalid names: Doryaspidinae Heintz 1967 [empty], Pteraspidinae Claypole 1885 [empty]
Subor. †Pteraspidoidei Janvier 1996
+
G. †Alaeckaspis Voichyshyn 1999
+
†Alaeckaspis ustetchkiensis Voichyshyn 1999
†Alaeckaspis verbivciensi Voichyshyn 1999
G. †Blieckaspis Elliott and Ilyes 1996
+
†Blieckaspis priscillae Elliott and Ilyes 1996
G. †Djurinaspis Novitskaya 1983
+
†Djurinaspis prima Novitskaya 1983
†Djurinaspis secunda Voichyshyn 2011
Fm. †Doryaspidae Randle and Sansom 2017
+
G. †Doryaspis White 1935
+
†Doryaspis arctica Pernè€gre 2003
†Doryaspis groenhorgensis Pernegre 2005
†Doryaspis nathorsti Lankester 1884
+
Invalid names: Scaphaspis nathorsti Lankester 1884 [replaced]
G. †Woodfjordaspis Pernègre 2006
+
†Woodfjordaspis felixi Pernegre 2006
G. †Xylaspis Pernegre 2003
+
†Xylaspis prima Pernegre 2003
G. †Eucyclaspis Denison 1970
+
†Eucyclaspis erroli Denison 1970
G. †Europrotaspis White 1961
+
G. †Lamiaspis Ilyes and Elliott 1994
+
†Lamiaspis longiripa Ilyes and Elliott 1994
G. †Larnovaspis Blieck 1984
+
†Larnovaspis brotzeni Brotzen 1933
†Larnovaspis goujeti Blieck 1984
†Larnovaspis kneri Lankester 1868
†Larnovaspis major Zych 1927
†Larnovaspis mogielnensis Blieck 1984
G. †Mylopteraspidella Blieck 1984
+
G. †Oreaspis Denison 1970
+
†Oreaspis ampla Bryant 1933
+
Invalid names: Protaspis nanus Bryant 1933 [synonym], Protaspis perlatus Bryant 1933 [synonym], Protaspis perryi Bryant 1933 [synonym]
†Oreaspis dunklei Denison 1970
†Oreaspis williamsi Denison 1970
G. †Panamintaspis Elliott and Ilyes 1996
+
†Panamintaspis snowi Elliott and Ilyes 1996
G. †Parapteraspis Stensio 1958
+
†Parapteraspis gracilis Stensio 1958
†Parapteraspis jackana White 1935
†Parapteraspis plana Stensio 1958
G. †Pirumaspis Ilyes and Elliott 1994
+
†Pirumaspis lancasteri Ilyes and Elliott 1994
G. †Podolaspis Zych 1931
+
Subg. †Podolaspis (Althaspis) Zych 1931
†Podolaspis danieli Voichyshyn 2011
†Podolaspis lerichei Zych 1927
†Podolaspis podolica Alth 1874
†Podolaspis zychi Brotzen 1933
+
G. †Clavulaspis Elliott et al. 2020
+
†Clavulaspis finis Elliott et al. 2020
Invalid names: Helaspis Elliott et al. 2000 [replaced]
G. †Cosmaspis Denison 1970
+
†Cosmaspis transversa Denison 1970
G. †Cyrtaspidichthys Whitley 1940
+
†Cyrtaspidichthys ovata Bryant 1932
†Cyrtaspidichthys sculpta Bryant 1933
G. †Lampraspis Denison 1970
+
†Lampraspis carmani Denison 1960
†Lampraspis tuberculata Denison 1970
G. †Psephaspis Orvig 1961
+
†Psephaspis bystrowi Halstead Tarlo 1964
†Psephaspis williamsi Orvig 1961
G. †Tuberculaspis Ilyes and Elliott 1994
+
†Tuberculaspis elyensis Ilyes and Elliott 1994
Fm. †Psammosteidae Traquair 1896
+
G. †Drepanaspis Schluter 1887
+
†Drepanaspis carteri McCoy 1851
†Drepanaspis edwardsi Halstead Tarlo 1964
†Drepanaspis gemuendensis Schluter 1887
†Drepanaspis jaegeri Halstead Tarlo 1964
†Drepanaspis schrieli Gross 1933
G. †Dyptychosteus Preobrazhenskii 1910
G. †Ganosteus Rohon 1901
+
†Ganosteus artus Mark-Kurik 1965
†Ganosteus stellatus Rohon 1901
G. †Oredezhosteus Moloshnikov 2009
+
†Oredezhosteus kuleshovi Moloshnikov 2009
G. †Placosteus Agassiz 1843
+
†Placosteus alatus Mark-Kurik 1965
†Placosteus undulatus Agassiz 1844
Subfm. †Psammosteinae Traquair 1896
+
G. †Elgaia Glinskiy 2017
+
†Elgaia luhai Glinskiy 2017
G. †Traquairosteus Halstead Tarlo 1964
+
†Traquairosteus falcatus Glinskiy and Nilov 2017
†Traquairosteus kiaeri Glinskiy and Nilov 2017
+
Invalid names: Psammosteus kiaeri Halstead Tarlo 1964 [replaced]
†Traquairosteus pectinatus Glinskiy and Nilov 2017
†Traquairosteus pustulatus Traquair 1897
†Traquairosteus ramosus Glinskiy and Nilov 2017
G. †Psammosteus Agassiz 1844
+
†Psammosteus arenatus Agassiz 1845
†Psammosteus asper Obruchev 1965
†Psammosteus bergi Mark-Kurik 1965
†Psammosteus cuneatus Obruchev 1965
†Psammosteus falcatus Gross 1942
†Psammosteus granulatus M'Coy 1848
†Psammosteus levis Obruchev 1965
†Psammosteus livonicus Obruchev 1965
†Psammosteus maeandrinus Agassiz 1845
†Psammosteus megalopteryx Trautschold 1880
†Psammosteus paradoxus Agassiz 1845
†Psammosteus pectinatus Obruchev 1965
†Psammosteus praecursor Obruchev 1947
†Psammosteus tchernovi Obruchev 1965
†Psammosteus tenuis Obruchev 1965
†Psammosteus waltergrossi Halstead Tarlo 1964
G. †Pycnosteus Preobrajenski 1911
+
†Pycnosteus obruchevi Halstead Tarlo 1964
†Pycnosteus palaeformis Preobrazhensky 1911
†Pycnosteus pauli Mark-Kurik 1965
G. †Vladimirolepis Glinskiy 2017
+
†Vladimirolepis proia Glinskiy 2017
G. †Pteraspis Kner 1847
+
Subg. †Pteraspis (Pteraspis) White 1956
†Pteraspis crouchi Lankester 1865
†Pteraspis rostrata Agassiz 1835
+
Invalid names: Plesiopteraspis Stensio 1958 [synonym]
Fm. †Rhinopteraspididae Novitskaya 2004
+
G. †Althaspis Zych 1931
+
†Althaspis leachi White 1938
†Althaspis senniensis Loeffler and Thomas 1980
†Althaspis tarloi Voichyshyn 2011
†Althaspis vimiensis White 1960
Invalid names: Brotzenaspis Stensio 1958 [synonym], Cymripteraspis White 1960 [synonym], Pseudopteraspis Stensio 1958 [synonym]
G. †Brachipteraspis Brotzen 1936
+
†Brachipteraspis latissima Zych in Pauca 1938
G. †Errivaspis Blieck 1984
+
†Errivaspis depressa Stensio 1958
†Errivaspis magnipinealis Brotzen 1933
†Errivaspis waynensis Blieck 1984
G. †Pavloaspis Voichyshyn 1999
+
†Pavloaspis pasternaki Voichyshyn 1999
G. †Rhinopteraspis Blieck 1984
+
†Rhinopteraspis crouchi Lankester 1868
†Rhinopteraspis dunensis Jaekel 1919
+
Invalid names: Pteraspis dunensis Roemer 1855 [replaced]
G. †Semipodolaspis Voichyshyn 2011
+
†Semipodolaspis slobodensis Voichyshyn 2011
G. †Unarkaspis Elliott 1983
+
†Unarkaspis schultzei Elliott 1983
Invalid names: Podolaspinae Zych 1931 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: marinesubp
Locomotion: actively mobilep
Created: 2017-04-17 15:11:50
Modified: 2017-04-17 15:11:50
Source: subp = subphylum, p = phylum
References: Kiessling 2004, Carroll 1988

Age range: base of the Ludlow to the top of the Pennsylvanian or 426.70000 to 298.90000 Ma

Collections (179 total)


Time interval Ma Country or state Original ID and collection number
Wenlock - Ludlow432.9 - 422.7Greenland Pteraspidiformes indet. (239394)
Ludlow426.7 - 422.7United Kingdom Kallostrakon alleni (239535) Kallostrakon macanuffi, Kallostrakon grindrodi, Kallostrakon podura (239534)
Ludlow - Pridoli426.7 - 419.62Canada (Nunavut) Ulutitaspis notidana (237776 237777) Ulutitaspis truncata (237773)
Early/Lower Lochkovian419.62 - 416.8Canada (Nunavut) Protopteraspis sp. indet. (238861)
Early/Lower Lochkovian419.62 - 416.8United Kingdom Protopteraspis gosseleti (238749 238779 238780 238781 238782 238783 238784 238785 238786 238788 238789 238790 238791 238792 238793 238794 238795 238797) Pteraspis rostrata (238750) Rhinopteraspis crouchi (238751)
Lochkovian - Early/Lower Lochkovian419.62 - 416.8Canada (Northwest Territories) Canadapteraspis thymostomata, Canadapteraspis helostomata (237849) Pteraspidae indet. (237847)
Gedinnine419.62 - 413.02Germany Pteraspis rostrata, Pteraspis jackana, Pteraspis traquairi, Rhinopteraspis crouchi (238362)
Lochkovian419.62 - 413.02Canada (Nunavut) Anchipteraspis crenulata (237786) Mitraspis cracens (238568) Pteraspididae indet. (14026) Rhachiaspis pteriga (237782 237785) Stegobranchiaspis baringensis (238034 238035) Stegobranchiaspis baringensis, Escharaspis alata (238128) Stegobranchiaspis baringensis, Unarkaspis schultzei (90322) Ulutitaspis notidana (237784) Ulutitaspis notidana, Rhachiaspis pteriga (237783) Unarkaspis schultzei (238129)
Lochkovian419.62 - 413.02France Althaspis vimiensis (239098) Protopteraspis gosseleti (26983)
Lochkovian419.62 - 413.02Canada (Northwest Territories) Canadapteraspis alocostomata (237813)
Lochkovian419.62 - 413.02Norway Canadapteraspis sp., Protopteraspis sp., Miltaspis anatirostrata (237820) Miltaspis anatirostrata, Larnovaspis goujeti, Protopteraspis vogti (237822) Protopteraspis primaeva (237816 237818) Protopteraspis sp. (26929 237815 237817 237821) Protopteraspis sp., Doryaspis sp. (237823 237826)
Lochkovian419.62 - 413.02Svalbard and Jan Mayen Protopteraspis micra (239099)
Lochkovian419.62 - 413.02USA (Wyoming) Protaspis transversa, Lampraspis tuberculata, Protaspis mcgrewi, Protaspis brevispina (153534)
Lochkovian419.62 - 413.02Russian Federation Pteraspidiformes indet., Protopteraspis sp., Unarkaspis sp., Pteraspididae indet. (90330)
Lochkovian419.62 - 413.02Ukraine Djurinaspis secunda (239217) Errivaspis depressa, Mylopteraspidella gracilis (238127) Larnovaspis brotzeni (239236) Larnovaspis goujeti (239250) Larnovaspis kneri, Larnovaspis brotzeni, Podolaspis zychi (239220) Larnovaspis kneri, Larnovaspis major (239235) Mylopteraspidella gracilis, Althaspis tarloi (239254) Podolaspis lerichei (239221) Podolaspis lerichei, Djurinaspis prima, Mylopteraspidella gracilis, Zascinaspis heintzi (239228) Podolaspis lerichei, Larnovaspis kneri, Larnovaspis major, Podolaspis podolica, Podolaspis zychi (239230) Podolaspis lerichei, Mylopteraspidella gracilis, Zascinaspis heintzi, Larnovaspis kneri, Podolaspis podolica (239231) Podolaspis lerichei, Zascinaspis heintzi, Larnovaspis major (239222) Podolaspis podolica (239252) Podolaspis sp., Larnovaspis sp. (26978 236556) Pteraspis major, Pteraspis rostrata, Larnovaspis kneri, Larnovaspis major (90349) Pteraspis podolicus, Pteraspis kneri, Pteraspididae indet. A, Pteraspididae indet. B, Pteraspididae indet. E, Pteraspis rostrata (90350) Semipodolaspis slobodensis (238873) Zascinaspis heintzi (239225 239227) Zascinaspis heintzi, Dnestraspis firma, Larnovaspis major, Larnovaspis mogielnensis, Parapteraspis plana (239251)
Lochkovian419.62 - 413.02United Kingdom (England) Althaspis sp. (90351) Europrotaspis crenulata, Althaspis leachi (90352) Pteraspis dairydinglensis (238454) Pteraspis rostrata (10679) Pteraspis sp. (26918)
Lochkovian419.62 - 413.02Canada (Nova Scotia) Protopteraspis sp. (14201)
Lochkovian419.62 - 413.02United Kingdom (Herefordshire) Parapteraspis jackana (239157) Pteraspis crouchi, Errivaspis waynensis, Pteraspis rostrata, Pteraspis rostrata, Pteraspis rostrata, Parapteraspis jackana (201736)
Lochkovian - Pragian419.62 - 410.62Ukraine Brachipteraspis latissima (239248) Larnovaspis major (239239) Podolaspis lerichei (239240) Pteraspis sp., Brotzenaspis kiaeri, Errivaspis magnipinealis (23447) Zascinaspis heintzi (239238)
Lochkovian - Pragian419.62 - 410.62USA (Ohio) Zascinaspis sp., Lampraspis carmani (26874)
Early/Lower Devonian419.62 - 393.47USA (Wyoming) Protaspis transversa, Lampraspis tuberculata, Protaspis mcgrewi, Protaspis brevispina (26179)
Lochkovian - Eifelian419.62 - 387.95Ukraine (Ternopil) Podolaspis lerichei, Alaeckaspis ustetchkiensis, Pavloaspis pasternaki, Djurinaspis prima, Mylopteraspidella gracilis, Zascinaspis heintzi, Podolaspis danieli, Dnestraspis firma, Larnovaspis major, Parapteraspis djurinensis, Parapteraspis plana, Podolaspis podolica, Podolaspis zychi (237769)
Late/Upper Lochkovian414.6 - 413.02Belgium Rhinopteraspis crouchi (238858)
Late/Upper Lochkovian - Pragian414.6 - 410.62Svalbard and Jan Mayen Xylaspis prima (238754)
Pragian413.02 - 410.62Svalbard and Jan Mayen Doryaspis nathorsti (238091 238092)
Pragian413.02 - 410.62Ukraine Brachipteraspis latissima (239255 239256 239259 239260) Brachipteraspis latissima, Larnovaspis sp. (239263) Mylopteraspidella gracilis (239247) Pteraspis major, Pteraspididae indet. D, Pteraspididae indet. F, Pteraspididae indet. G, Podolaspis lerichei, Brachipteraspis latissima (90348)
Pragian413.02 - 410.62Russian Federation Protopteraspididae indet., Protopteraspididae indet. 1, Pteraspidiformes indet., Pteraspididae indet., Miltaspis sp., Gigantaspis sp. (90331)
Pragian413.02 - 410.62United Kingdom Althaspis senniensis (238752)
Siegen413.02 - 410.62Germany (North Rhine-Westphalia) Rhinopteraspis dunensis, Drepanaspis schrieli (232143)
Pragian - Emsian413.02 - 393.47Germany Rhinopteraspis dunensis (239017)
Pragian - Emsian413.02 - 393.47Svalbard and Jan Mayen Doryaspis arctica, Woodfjordaspis felixi (238095) Doryaspis nathorsti, Doryaspis arctica (237853) Gigantaspis isachseni (237852 238272) Gigantaspis isachseni, Gigantaspis minima (237851) Gigantaspis laticephala (238274)
Pragian - Emsian413.02 - 393.47Norway Doryaspis arctica, Doryaspis groenhorgensis, Zascinaspis laticephala (233178) Doryaspis groenhorgensis (239307)
Early/Lower Emsian410.62 - 399.5Germany (Gemuenden) Drepanaspis gemuendensis (26903)
Early/Lower Emsian410.62 - 399.5Germany Drepanaspis gemuendensis (230899) Rhinopteraspis dunensis (238361)
Early/Lower Emsian410.62 - 399.5Germany (Alken) Drepanaspis sp., Rhinopteraspis dunensis (27049)
Emsian410.62 - 393.47USA (Wyoming) Protaspis bucheri, Protaspis dorfi, Protaspis ovata, Protaspis sculpta, Oreaspis dunklei, Oreaspis williamsi, Oreaspis ampla (26176)
Emsian410.62 - 393.47USA (Utah) Blieckaspis priscillae (233157) Psephaspis williamsi (233171)
Emsian410.62 - 393.47Belarus (Vitebsk) Psammosteoidei indet. (232875)
Emsian410.62 - 393.47Belarus (Gormel) Psammosteus toriensis, Psammosteoidei indet. (232955)
Late/Upper Emsian399.5 - 393.47USA (Nevada) Pteraspidinae indet., Pirumaspis lancasteri, Tuberculaspis elyensis, Lamiaspis longiripa (237780)
Late/Upper Emsian399.5 - 393.47USA (California) Pteraspididae indet., Blieckaspis priscillae (206448) Pteraspididae indet., Panamintaspis snowi (206449)
Eifelian393.47 - 387.95Belarus Pycnosteus sp. (225881 225882)
Eifelian393.47 - 387.95Brazil (Para) Pteraspis sp. (37137)
Eifelian393.47 - 387.95Belarus (Vitebsk) Pteraspidiformes indet., Psammosteoidei indet. (232958)
Eifelian393.47 - 387.95Russian Federation Pycnosteus sp., Protopteraspididae indet 2, Psammosteus sp., Psammosteidae indet. (90332)
Eifelian393.47 - 387.95USA (Utah) Eucyclaspis erroli (238263)
Eifelian393.47 - 387.95USA (Idaho) Scutellaspis wilsoni, Ecphymaspis idahoensis (238356)
Eifelian393.47 - 387.95Estonia Pycnosteus sp., Ganosteus sp. (225793)
Early/Lower Givetian387.95 - 385.9Russian Federation Psammosteus sp., Psammosteidae indet. (90333)
Early/Lower Givetian387.95 - 385.9Estonia Pycnosteus tuberculatus, Psammosteus bergi (225851) Vladimirolepis proia, Ganosteus artus, Pycnosteus palaeformis (225804) Vladimirolepis proia, Ganosteus artus, Pycnosteus pauli (225803)
Early/Lower Givetian387.95 - 385.9Latvia Vladimirolepis proia, Ganosteus artus, Pycnosteus palaeformis (225850) Vladimirolepis proia, Ganosteus artus, Pycnosteus pauli, Pycnosteus palaeformis (225802)
Early/Lower Givetian387.95 - 385.9Belarus Pycnosteus sp. (225886)
Givetian387.95 - 382.31Latvia Psammosteus livonicus, Placosteus undulatus (239144)
Givetian387.95 - 382.31Estonia Elgaia luhai (239669) Ganosteus stellatus, Vladimirolepis proia, Ganosteus artus, Pycnosteus pauli, Pycnosteus palaeformis (239664) Placosteus alatus (239670) Psammosteus bergi (239672)
Givetian387.95 - 382.31Svalbard and Jan Mayen (Spitsbergen) Psammosteidae indet., Psammosteus arenatus (225592)
Givetian387.95 - 382.31Russian Federation Psammosteus waltergrossi, Traquairosteus falcatus (239668)
Givetian387.95 - 382.31Canada (Alberta) Clavulaspis finis (238355)
Givetian387.95 - 382.31Belarus (Gormel) Pycnosteus sp., Psammosteus sp., Ganosteus sp., Ganosteus stellatus (242242)
Givetian387.95 - 382.31Belarus (Brest) Ganosteus sp., Psammosteoidei indet. (242243) Pycnosteus sp., Ganosteus sp. (242244)
Late/Upper Givetian383.4 - 382.31Russian Federation Psammosteus sp. (90334)
Late/Upper Givetian383.4 - 382.31Belarus Psammosteus praecursor (225888)
Late/Upper Givetian383.4 - 382.31Latvia Placosteus undulatus, Traquairosteus ramosus (239671)
Late/Upper Givetian383.4 - 382.31Estonia Psammosteus spp., Ganosteus stellatus (225778)
Early/Lower Frasnian382.31 - 379.0Russian Federation Oredezhosteus kuleshovi (90346) Psammosteus praecursor (90343) Psammosteus sp. (90335)
Early/Lower Frasnian382.31 - 379.0Latvia Oredezhosteus livonicus (90347)
Frasnian382.31 - 372.15Canada (Nunavut) Traquairosteus kiaeri (239676)
Frasnian382.31 - 372.15United Kingdom Psammosteidae indet. (239215)
Frasnian382.31 - 372.15Russian Federation Psammosteus cuneatus (239673) Psammosteus levis, Traquairosteus ramosus, Traquairosteus pectinatus (239674) Psammosteus maeandrinus, Psammosteus asper, Traquairosteus pectinatus (239134) Psammosteus tenuis (239675)
Middle Frasnian379.0 - 375.2Russian Federation (Saint Petersburg) Psammosteus megalopteryx (196703)
Middle Frasnian379.0 - 375.2Canada (Nunavut) Perscheia pulla (236499)
Middle Frasnian - Late/Upper Frasnian379.0 - 372.15United Kingdom Psammosteus megalopteryx (239413)
Late/Upper Frasnian375.2 - 372.15Russian Federation Obruchevia heckeri, Psammosteus falcatus (238745) Psammosteus sp. (90336)
Late/Upper Frasnian375.2 - 372.15United Kingdom (Scotland) Traquairosteus pustulatus, Traquairosteus falcatus (176215)
Famennian372.15 - 358.86Greenland Psammosteus sp. (238738)
Pennsylvanian323.4 - 298.9Ireland Psammosteus granulatus (132818)
Pennsylvanian323.4 - 298.9United Kingdom Psammosteus vermicularis (132819)