| Basic info | Taxonomic history | Classification | Included Taxa |
| Morphology | Ecology and taphonomy | External Literature Search | Age range and collections |
Lioharpes
Taxonomy
Lioharpes was named by Whittington (1950). Its type is Harpes venulosus. It was considered monophyletic by Johnson (2024).
It was assigned to Ptychopariida by Sepkoski (2002); and to Harpetidae by Ebach and McNamara (2002), Jell and Adrain (2002), Beech and Lamsdell (2021), Johnson (2024).
It was assigned to Ptychopariida by Sepkoski (2002); and to Harpetidae by Ebach and McNamara (2002), Jell and Adrain (2002), Beech and Lamsdell (2021), Johnson (2024).
Species
Species lacking formal opinion data
Synonymy list
| Year | Name and author |
|---|---|
| 1950 | Lioharpes Whittington |
| 2002 | Lioharpes Ebach and McNamara |
| 2002 | Lioharpes Jell and Adrain p. 398 |
| 2002 | Lioharpes Sepkoski |
| 2021 | Lioharpes Beech and Lamsdell p. 2221 |
| 2024 | Lioharpes Johnson p. 54 |
Is something missing? Join the Paleobiology Database and enter the data
|
|
If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Lioharpes Whittington 1950
show all | hide all
Subg. †Lioharpes (Lioharpes) Pribyl and Vanek 1986
†Lioharpes altaicus Weber 1932
†Lioharpes ammari Johnson 2024
†Lioharpes bischofi Roemer 1852
†Lioharpes crassimargo Novak 1890
†Lioharpes galea Johnson 2024
†Lioharpes hastatus Lütke 1965
†Lioharpes morocconensis Johnson 2024
†Lioharpes polaris Maksimova 1977
†Lioharpes saredra Johnson 2024
†Lioharpes scopulum Johnson 2024
†Lioharpes sculptus Hawle and Corda 1847
†Lioharpes vektori Doubrava 1991
†Lioharpes venetus Gortani 1915
†Lioharpes venulosus Whittington 1950
hide
Invalid names: Harpes ruderalis Hawle and Corda 1847 [synonym], Lioharpes (Lioharpes) venulosus Pribyl and Vanek 1986 [synonym], Lioharpes klukovicensis Vanek et al. 1992 [synonym]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| R. G. Johnson 2024 | Cephalon ovoid. Occipital furrow broad (sag.), equal to or wider (exsag.) than posterior
border. Width (tr.) of posterior border between 31% to 40% of occipital ring width (tr.). Genal area moderately broad, inner margin of fringe across anterior of genal area straight and Eye lobes with three lenses each. Tubercles on occipital ring, palpebral lobes and dorsal surface of internal rim. Girder narrow (sag. And exsag.), 1.5% to 2.2% of genicranium width (tr.). Brim width ratio between 0.70 and 0.85. Extension of girder meeting internal rim halfway down prolongation. Thorax with 20 to 24 thoracic segments. Pleural furrows shallow and narrow, and distal ends of pleurae rounded. |
Measurements
No measurements are available
|
|
||||
|
|
||||
|
|
||||
|
|
||||
|
|
||||
| Source: g = genus, f = family, c = class | |||||
| References: Aberhan et al. 2004, Whittington et al. 1997, Jell and Adrain 2002 | |||||
Age range: base of the Lochkovian to the top of the Eifelian or 419.62000 to 387.95000 Ma
Collections (27 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Lochkovian | Czechia | L. ruderalis (10315) L. sp. (14119 14121) | |
| Lochkovian | Russian Federation (Sverdlovskaya Oblast) | Harpes venulosus (81749) | |
| Lochkovian | Uzbekistan | L. venulosus (181368 181369) | |
| Early/Lower Devonian | Czechia (Barrandia) | L. crassimargo (12183) | |
| Early/Lower Devonian - Eifelian | Czechia | L. perneri (12148) | |
| Pragian | Czechia (Praha) | L. sculptus (157332) L. venulosus (229099) | |
| Pragian | Czechia | L. venulosus (10342) | |
| Pragian | Russian Federation (Sverdlovskaya Oblast) | Harpes venulosus (81793) | |
| Pragian | Czechia (Prague) | L. venulosus (12071) | |
| Pragian - Early/Lower Emsian | Russian Federation (Sverdlovskaya Oblast) | Harpes venulosus (81797) | |
| Pragian - Early/Lower Emsian | Italy | Harpes venetus (26245 26248) | |
| Pragian - Middle Devonian | Czechia | L. venulosus (10344) | |
| Early/Lower Emsian | Russian Federation (Arkhangel'skaya Oblast') | Harpes polaris (40703) | |
| Early/Lower Emsian | Czechia | L. crassimargo (229101) | |
| Early/Lower Emsian - Late/Upper Emsian | Czechia | L. crassimargo (10462) | |
| Zlichovian | Czechia (Barrandia) | L. crassimargo (12112 12143) | |
| Zlichovian | Czechia | L. crassimargo (10460 229100) L. sp. (10467) | |
| Eifelian | Czechia (Koneprusy) | L. montageni (12257) | |
| Eifelian | Czechia (Barrandia) | L. montageni (12252) |