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Varunidae

Malacostraca - Decapoda - Varunidae

Taxonomy
Varunidae was named by Milne-Edwards (1853). It is extant. Its type is Varuna.

It was assigned to Grapsoidea by Schweitzer et al. (2010), Schweitzer et al. (2023).

Synonymy list
YearName and author
1853Varunidae Milne-Edwards
2010Varunidae Schweitzer et al. p. 142
2023Varunidae Schweitzer et al. p. 5

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—Hymenostraca(Rolfe 1969)
superclassMulticrustaceaRegier et al. 2010
classMalacostraca()
RankNameAuthor
subclassEumalacostracaGrobben 1892
—CaridoidaHessler 1982
orderDecapoda
suborderPleocyemataBurkenroad 1963
—Reptantia(Boas)
—MeiuraSaint Laurent 1980
infraorderBrachyura()
—Eubrachyurade Saint Laurent 1980
—ThoracotremataGuinot 1977
superfamilyGrapsoideaMacLeay 1838
familyVarunidaeMilne-Edwards 1853
familyVarunidaeMilne-Edwards 1853

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Varunidae Milne-Edwards 1853
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Subfm. Asthenognathinae Stimpson 1858
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G. Asthenognathus Stimpson 1858
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†Asthenognathus cornishorum Schweitzer and Feldmann 1999
†Asthenognathus globosa Karasawa 1990
†Asthenognathus hexagonum Rathbun 1909
†Asthenognathus inaequipes Stimpson 1858
†Asthenognathus laverdensis De Angeli and Garassino 2006
†Asthenognathus microspinus Casadio et al. 2004
†Asthenognathus rakosensis Müller 2006
†Asthenognathus urretae Schweitzer and Feldmann 2001
G. †Globihexapus Schweitzer and Feldmann 2001
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†Globihexapus kosekii Kato 2005
†Globihexapus paxillus Schweitzer and Feldmann 2001
Subfm. Cyclograpsinae Milne Edwards 1853
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G. Austrohelice Sakai et al. 2006
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†Austrohelice manneringi Feldmann et al. 2008
G. Cyclograpsus Milne-Edwards 1937
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†Cyclograpsus directus Karasawa 1989
†Cyclograpsus lwangi Hu and Tao 1999
†Cyclograpsus rectangularis Karasawa 1989
G. Helice De Haan 1833
G. †Miosesarma Karasawa 1989
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†Miosesarma japonica Karasawa 1989
†Miosesarma naguraense Kato 1996
Subfm. Gaeticinae Davie and Ng 2007
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G. Sestrostoma Davie and Ng 2007
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†Sestrostoma hichiro Karasawa and Tanaka 2005
G. †Sakakurapsus Karasawa 2018
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†Sakakurapsus kogisorum Karasawa 2018
Subfm. Thalassograpsinae Davie and Ng 2007
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G. Thalassograpsus Tweedie 1950
Subfm. Varuninae Milne-Edwards 1853
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G. Acmaeopleura Stimpson 1858
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Acmaeopleura rotunda Rathbun 1909
G. Brachynotus De Haan 1833
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†Brachynotus corallinus Beschin et al. 2007
†Brachynotus februarius Müller 1974
†Brachynotus oligocenicus De Angeli et al. 2010
G. Eriocheir De Haan 1835
+
Eriocheir japonica De Haan 1835
G. Hemigrapsus Dana 1851
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Hemigrapsus nudus Dana 1851 [purple shore crab]
†Hemigrapsus oregonensis Dana 1851 [yellow shore crab]
G. Utica White 1847
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Utica haswelli Wintle 1886
Utica yarraensis Wintle 1886
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Varuna angustifrons Karasawa 1993
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Locomotion: actively mobilec
Life habit: infaunalsuperf
Diet: detritivoresuperf
Vision: well-developedc
Created: 2022-08-23 13:46:39
Modified: 2022-09-18 00:50:49
Source: superf = superfamily, c = class
References: Aberhan 1992, Valdez and Clapham 2023

Age range

Maximum range based only on fossils: base of the Ypresian to the top of the Holocene or 56.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 48.07 Ma

Collections (39 total)


Time interval Ma Country or state Original ID and collection number
Ypresian56.0 - 48.07Italy Brachynotus corallinus (140158 207470 207474)
Lutetian48.07 - 41.03Argentina (Santa Cruz) Asthenognathus urretae (202316)
Priabonian37.71 - 33.9Italy Brachynotus oligocenicus (207550)
Rupelian33.9 - 27.3Japan (Saga) Varuna angustifrons (127692 202440)
Rupelian33.9 - 27.3Italy (Vicenza) Asthenognathus laverdensis (127496) Brachynotus oligocenicus (207617)
Chattian27.3 - 23.04USA (Washington) Asthenognathus cornishorum (221302 221303 221304)
Early/Lower Miocene23.04 - 15.98Argentina (Rio Negro) Asthenognathus microspinus (45689)
Burdigalian20.45 - 15.98Argentina Asthenognathus urretae (43002)
Burdigalian20.45 - 15.98USA (Oregon) Globihexapus paxillus (202317)
Langhian15.98 - 13.82Japan (Okayama) Miosesarma japonicum (127702 127706 127707)
Langhian15.98 - 13.82Japan (Shimane) Miosesarma japonicum (202462)
Middle Miocene15.98 - 11.63USA (Oregon) Globihexapus paxillu (202348)
Middle Miocene15.98 - 11.63Japan (Toyama) Miosesarma japonicum (202474 202475)
Badenian13.82 - 12.8Hungary Asthenognathus rakosensis (149366) Brachynotus februarius (138128 138812 164315 207750 207752 207753)
Pliocene5.333 - 2.58Italy (Grosseto) Asthenognathus sp. (221599)
Pliocene - Holocene5.333 - 0.0USA (California) Hemigrapsus sp. (138983)
Piacenzian3.6 - 2.58New Zealand Austrohelice manneringi (79321)
Pleistocene2.58 - 0.0117USA (California) Hemigrapsus nudus (138720) Hemigrapsus nudus, Hemigrapsus oregonensis (138719)
Pleistocene - Holocene2.58 - 0.0USA (California) Hemigrapsus nudus (138721) Hemigrapsus oregonensis (138724)
Middle Pleistocene0.774 - 0.129USA (California) Hemigrapsus nudus, Hemigrapsus oregonensis (138723)
Late/Upper Pleistocene0.129 - 0.0117USA (California) Hemigrapsus nudus, Hemigrapsus oregonensis (160361)
Holocene0.0117 - 0.0USA (Florida) Cyclograpsus integer (146566)