PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Compsoscorpius buthiformis

Arachnida - Scorpiones

Taxonomy

Anthracoscorpio buthiformis was named by Pocock (1911). It is a compression fossil. Its type locality is Sparth Bottoms, Rochdale (BMNH coll), which is in a Westphalian A delta plain shale in the United Kingdom.

It was recombined as Eoscorpius buthiformis by Petrunkevitch (1949); it was recombined as Buthiscorpius buthiformis by Petrunkevitch (1953), Wills (1960) and Kjellesvig-Waering (1986); it was recombined as Compsoscorpius buthiformis by Legg et al. (2012) and Dunlop et al. (2013).

Synonyms

Buthiscorpius major was named by Wills (1960). Its type specimen is GSM Za 2926, an exoskeleton, and it is a compression fossil. Its type locality is Trowell Colliery, which is in a Westphalian B deltaic shale in the United Kingdom.

It was recombined as Allobuthiscorpius major by Kjellesvig-Waering (1986); it was synonymized subjectively with Compsoscorpius buthiformis by Legg et al. (2012) and Dunlop et al. (2013).
Lichnoscorpius minutus was named by Petrunkevitch (1949). Its type specimen is BMNH In. 31266 and is a compression fossil. Its type locality is Coseley, near Dudley (Egginton coll), which is in a Westphalian B interdistributary bay shale in the United Kingdom. It is the type species of Lichnoscorpius.

It was recombined as Anthracoscorpio minutus by Petrunkevitch (1953); it was synonymized subjectively with Compsoscorpius buthiformis by Legg et al. (2012) and Dunlop et al. (2013).
Allobuthus macrostethus was named by Kjellesvig-Waering (1986). Its type specimen is BU 720, an exoskeleton, and it is a compression fossil. Its type locality is Coseley, near Dudley (Birmingham University coll), which is in a Westphalian B interdistributary bay shale in the United Kingdom.

It was synonymized subjectively with Compsoscorpius buthiformis by Legg et al. (2012) and Dunlop et al. (2013).
Coseleyscorpio lanceolatus was named by Kjellesvig-Waering (1986). Its type specimen is BMNH In. 31262, an exoskeleton, and it is a compression fossil. Its type locality is Coseley, near Dudley (Egginton coll), which is in a Westphalian B interdistributary bay shale in the United Kingdom.

It was synonymized subjectively with Compsoscorpius buthiformis by Legg et al. (2012) and Dunlop et al. (2013).
Leioscorpio pseudobuthiformis was named by Kjellesvig-Waering (1986). Its type specimen is BMNH In. 22832 and is an impression. Its type locality is Coseley, near Dudley (Hind coll), which is in a Westphalian B interdistributary bay shale in the United Kingdom.

It was synonymized subjectively with Compsoscorpius buthiformis by Legg et al. (2012) and Dunlop et al. (2013).
Pseudobuthiscorpius labiosus was named by Kjellesvig-Waering (1986). Its type specimen is BMNH I.1555 and is a compression fossil. Its type locality is Coseley, near Dudley (Johnson coll), which is in a Westphalian B interdistributary bay shale in the United Kingdom.

It was synonymized subjectively with Compsoscorpius buthiformis by Legg et al. (2012) and Dunlop et al. (2013).
Compsoscorpius elegans was named by Petrunkevitch (1949). Its type specimen is BMNH I.7883 and is a compression fossil. Its type locality is Coseley, near Dudley (Madeley coll), which is in a Westphalian B interdistributary bay shale in the United Kingdom. It is the type species of Compsoscorpius.

It was synonymized subjectively with Compsoscorpius buthiformis by Legg et al. (2012) and Dunlop et al. (2013).
Compsoscorpius elongatus was named by Petrunkevitch (1949). Its type specimen is BMNH In. 15862 and is a compression fossil. Its type locality is Coseley, near Dudley (Johnson coll), which is in a Westphalian B interdistributary bay shale in the United Kingdom.

It was synonymized subjectively with Compsoscorpius elegans by Kjellesvig-Waering (1986) and Jeram (1994); it was synonymized subjectively with Compsoscorpius buthiformis by Legg et al. (2012) and Dunlop et al. (2013).
Typhlopisthacanthus anglicus was named by Petrunkevitch (1949). Its type specimen is BMNH In. 31261, an exoskeleton, and it is a compression fossil. Its type locality is Coseley, near Dudley (Egginton coll), which is in a Westphalian B interdistributary bay shale in the United Kingdom.

It was synonymized subjectively with Compsoscorpius elegans by Kjellesvig-Waering (1986) and Jeram (1994); it was synonymized subjectively with Compsoscorpius buthiformis by Legg et al. (2012) and Dunlop et al. (2013).

Synonymy list

Year	Name and author
1911	Anthracoscorpio buthiformis Pocock p. 24 figs. Pl 1, fig 2; pl 2, fig 1; text-figs 6-8
1949	Typhlopisthacanthus anglicus Petrunkevitch p. 145 figs. 143, 182
1949	Lichnoscorpius minutus Petrunkevitch p. 148 figs. 144, 145, 181
1949	Compsoscorpius elegans Petrunkevitch p. 149 figs. 152-154, 183-185
1949	Compsoscorpius elongatus Petrunkevitch p. 150 figs. 147-150, 186-188
1949	Eoscorpius buthiformis Petrunkevitch p. 153
1953	Anthracoscorpio minutus Petrunkevitch p. 30
1953	Buthiscorpius buthiformis Petrunkevitch p. 32 figs. 31-34, 122
1953	Compsoscorpius elegans Petrunkevitch p. 32
1953	Compsoscorpius elongatus Petrunkevitch p. 33
1953	Typhlopisthacanthus anglicus Petrunkevitch p. 34
1960	Buthiscorpius buthiformis Wills p. 277 figs. Pls 46, figs 1-5; pl 47, figs 1-6; pl 48, figs 1-7; text-figs 1-9
1960	Buthiscorpius major Wills p. 300 figs. Pl 51, figs 1-3; pl 52; text-figs 14-16
1986	Buthiscorpius buthiformis Kjellesvig-Waering p. 102 figs. 40, 110F
1986	Allobuthiscorpius major Kjellesvig-Waering p. 105
1986	Lichnoscorpius minutus Kjellesvig-Waering p. 110 fig. 45
1986	Allobuthus macrostethus Kjellesvig-Waering p. 112 figs. 110C, 113B4
1986	Coseleyscorpio lanceolatus Kjellesvig-Waering p. 113
1986	Leioscorpio pseudobuthiformis Kjellesvig-Waering p. 209 fig. 92
1986	Pseudobuthiscorpius labiosus Kjellesvig-Waering p. 219 figs. 97, 112J
1986	Compsoscorpius elegans Kjellesvig-Waering p. 236 figs. 105-107
1994	Compsoscorpius elegans Jeram p. 530
2012	Compsoscorpius buthiformis Legg et al. p. 5 figs. 3,5; videos 1,2
2013	Compsoscorpius buthiformis Dunlop et al. p. 35

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
subphylum	Chelicerata	Heymons 1901
—	Euchelicerata	Weygoldt and Paulus 1979
—	Prosomapoda

Rank	Name	Author
—	Planaterga
—	Dekatriata
—	Sclerophorata	Kamenz et al. 2011
class	Arachnida	Lamarck 1801
—	Stomothecata
order	Scorpiones	Latreille 1817
suborder	Orthosternina	()
superfamily	Scorpionoidea
genus	Compsoscorpius
species	buthiformis	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Compsoscorpius buthiformis Pocock 1911

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Invalid names: Allobuthus macrostethus Kjellesvig-Waering 1986 [synonym], Buthiscorpius major Wills 1960 [synonym], Compsoscorpius elegans Petrunkevitch 1949 [synonym], Compsoscorpius elongatus Petrunkevitch 1949 [synonym], Coseleyscorpio lanceolatus Kjellesvig-Waering 1986 [synonym], Leioscorpio pseudobuthiformis Kjellesvig-Waering 1986 [synonym], Lichnoscorpius minutus Petrunkevitch 1949 [synonym], Pseudobuthiscorpius labiosus Kjellesvig-Waering 1986 [synonym], Typhlopisthacanthus anglicus Petrunkevitch 1949 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Grouping:

solitaryc

Environment:

terrestrialc

Locomotion:

actively mobilec

Created:

2017-04-17 09:55:58

Modified:

2017-04-17 09:55:58

Source: c = class

Reference: Bush and Bambach 2015

Age range: base of the Westphalian A to the top of the Westphalian B or 318.70000 to 314.60000 Ma

Collections (16 total)

Time interval	Ma	Country or state	Original ID and collection number
Westphalian A	318.7 - 316.9	United Kingdom (Durham)	Compsoscorpius elegans (153598)
Westphalian A	318.7 - 316.9	United Kingdom (England)	Anthracoscorpio buthiformis (type locality: 34905) Compsoscorpius elegans (146853)
Westphalian A	318.7 - 316.9	United Kingdom (Lancashire)	Compsoscorpius elegans (153597 153599)
Westphalian B	316.9 - 314.6	United Kingdom (Yorkshire)	Compsoscorpius elegans (153600 153601 153602 153603 153604)
Westphalian B	316.9 - 314.6	United Kingdom (Nottinghamshire)	Buthiscorpius major (153539)
Westphalian B	316.9 - 314.6	United Kingdom (Staffordshire)	Anthracoscorpio buthiformis (34906 34908) Buthiscorpius buthiformis (153540) Lichnoscorpius minutus, Typhlopisthacanthus anglicus, Anthracoscorpio buthiformis (34907)
Westphalian B	316.9 - 314.6	United Kingdom (England)	Compsoscorpius elongatus, Anthracoscorpio buthiformis (125238)