PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Tylocidaris

Echinoidea - Cidaroida - Psychocidaridae

Taxonomy

Tylocidaris was named by Pomel (1883) [Sepkoski's age data: ]. Its type is Cidaris clavigera.

It was assigned to Cidaridae by Cooke (1941); to Psychocidaridae by Fell (1966); to Psychocidarinae by Gravesen (1993); and to Cidaroida by Sepkoski (2002).

Species

T. (Sardocidaris), T. (Tylocidaris), T. clavigera (syn. Cidaris heberti), T. gosae, T. salina, T. vexilifera, T. walcotti, T. windi

Species lacking formal opinion data

T. bermudez, T. bermudezi, T. bruennichi, T. hardouini, T. inexpectata, T. ramondi, T. sorigneti, T. trempinus, T. velifera, T. versiculosa, T. vesicolosa, T. vexillifera

Synonymy list

Year	Name and author
1883	Tylocidaris Pomel
1941	Tylocidaris Cooke p. 6
1966	Tylocidaris Fell p. U339
1993	Tylocidaris Gravesen p. 50
2002	Tylocidaris Sepkoski

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
—	Ambulacraria
phylum	Echinodermata
subphylum	Eleutherozoa

Rank	Name	Author
—	Echinozoa	()
class	Echinoidea	()
subclass	Cidaroidea	(Claus 1880)
order	Cidaroida	()
superfamily	Cidaridea	Gray 1825
family	Psychocidaridae
genus	Tylocidaris	Pomel 1883

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Tylocidaris Pomel 1883

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Subg. †Tylocidaris (Sardocidaris) Lambert 1907

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†Tylocidaris (Sardocidaris) scarabellii Stefanini 1907

Invalid names: Sardocidaris piae Lambert 1907 [synonym]

Subg. †Tylocidaris (Tylocidaris) Pomel 1883

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†Tylocidaris (Tylocidaris) abildgaardi Ravn 1928

†Tylocidaris (Tylocidaris) oedumi Brünnich Nielson 1938

†Tylocidaris (Tylocidaris) ravni Brotzen 1959

†Tylocidaris (Tylocidaris) rosenkrantzei Brotzen 1959

†Tylocidaris clavigera Mantell 1822

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Invalid names: Cidaris heberti Desor 1855 [synonym]

†Tylocidaris gosae Schluter 1892

†Tylocidaris salina Cooke 1959

†Tylocidaris vexilifera Schlüter 1982

†Tylocidaris walcotti Clark 1891

†Tylocidaris windi Brotzen 1959

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

high Mg calciteo

Ontogeny:

accretion, addition of partsp

Environment:

marineo

Locomotion:

slow-movingo

Life habit:

low-level epifaunalo

Diet:

grazero

Diet 2:

omnivoreo

Vision:

limitedc

Dispersal:

watero

Dispersal 2:

planktonico

Created:

2004-02-28 12:08:25

Modified:

2009-12-13 03:42:21

Source: o = order, c = class, p = phylum

References: Aberhan 1992, Kiessling 2004

Age range: base of the Late/Upper Barremian to the top of the Burdigalian or 125.77000 to 15.98000 Ma

Collections (47 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Barremian	125.77 - 121.4	France (Drôme)	T. sp. (75576)
Early/Lower Cenomanian - Late/Upper Albian	100.5 - 100.5	Spain	T. sorigneti (3832)
Early/Lower Cenomanian	100.5 - 93.9	United Kingdom (England)	T. velifera (59711)
Early/Lower Cenomanian	100.5 - 93.9	United Kingdom	T. velifera (59682 59688)
Late/Upper Cenomanian	100.5 - 93.9	Czechia	T. vesicolosa (90511) T. vesicolosa, T. sorigneti, T. versiculosa (90513)
Early/Lower Turonian	93.9 - 89.8	Czechia	T. vesicolosa, T. sorigneti, T. versiculosa (90516 90517 90518)
Early/Lower Turonian	93.9 - 89.8	France	T. clavigera (59620)
Turonian	93.9 - 89.8	Belgium	T. clavigera (155279)
Turonian	93.9 - 89.8	United Kingdom (England)	T. clavigera (38595)
Coniacian	89.8 - 85.7	United Kingdom (England)	T. clavigera (38597)
Early/Lower Santonian	85.7 - 83.6	Spain	T. clavigera (74423)
Early/Lower Santonian - Middle Santonian	85.7 - 83.6	Sweden	T. sp. (70316)
Middle Santonian	85.7 - 83.6	Germany	T. gosae (178204)
Late/Upper Maastrichtian	72.2 - 66.0	Netherlands (Limburg)	T. hardouini (32167)
Maastrichtian	72.2 - 66.0	Spain (Santander)	T. ramondi (157019 157021 157022 157023) T. sp. (157004) T. trempinus (157006 157008)
Late/Upper Maastrichtian - Danian	72.2 - 61.66	Netherlands	T. hardouini, T. inexpectata (32160)
Danian	66.0 - 61.66	Netherlands	T. hardouini (3122)
Danian	66.0 - 61.66	Netherlands (Limburg)	T. bruennichi, T. hardouini (32170)
Danian	66.0 - 61.66	Denmark	T. abildgaardi (78711) T. oedumi (35040) T. vexillifera (89866)
Late/Upper Paleocene	59.24 - 56.0	USA (New Jersey)	T. walcotti (155898 155902)
Thanetian	59.24 - 56.0	USA (New Jersey)	T. walcotti (3113 81775)
Thanetian	59.24 - 56.0	USA (Alabama)	T. salina (155900 155901)
Lutetian - Bartonian	48.07 - 37.71	Cuba (all)	T. bermudezi (8210)
Priabonian	37.71 - 33.9	Cuba (Pinar del Rio)	T. bermudez (8209)
Aquitanian	23.04 - 20.45	Italy	Sardocidaris piae (209212)
Aquitanian - Burdigalian	23.04 - 15.98	Italy (Sardinia)	T. sp. (187424 187425 187426)
Burdigalian	20.45 - 15.98	San Marino	T. scarabellii (209200)
Burdigalian - Langhian	20.45 - 13.82	Italy	Cidaris scarabellii (209211) T. scarabellii (209206 209209)