PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Hoplitoidea

Cephalopoda - Ammonoida

Taxonomy

Hoplitaceaea was named by Douvillé (1890). Its type is Hoplites.

It was corrected as Hoplitoidea by Kennedy (2005), Kennedy and Latil (2007), Aly et al. (2008), Kennedy et al. (2008), Kennedy et al. (2008), Mosavinia et al. (2014), Shigeta (2016), Kennedy and Gale (2016), Kennedy (2018), Tajika et al. (2018), El Qot (2018); it was corrected as Hoplitaceae by Wright (1952), Nagy (1970), Förster (1975), Wiedmann (1978), Cooper (1978), Cooper and Kennedy (1979), Summesberger (1979), Kennedy et al. (1981), Kennedy (1984), Klinger and Kennedy (1989), Kennedy and Simmons (1991), Kennedy (1995), Wright et al. (1996), Cecca (1997), Kennedy et al. (1997), Kennedy et al. (1998), Kennedy and Summesberger (1999), Szives (2007), Sharifi et al. (2016), Robert et al. (2018).

It was assigned to Ancyloceratina by Cooper (1978); to Ammonoidea by Kennedy and Simmons (1991); and to Ammonitina by Wright (1952), Nagy (1970), Förster (1975), Wiedmann (1978), Cooper and Kennedy (1979), Summesberger (1979), Kennedy et al. (1981), Kennedy (1984), Klinger and Kennedy (1989), Kennedy et al. (1992), Kennedy (1995), Wright et al. (1996), Cecca (1997), Kennedy et al. (1997), Kennedy et al. (1998), Kennedy and Summesberger (1999), Kennedy (2005), Szives (2007), Kennedy and Latil (2007), Aly et al. (2008), Kennedy et al. (2008), Kennedy et al. (2008), Mosavinia et al. (2014), Shigeta (2016), Kennedy and Gale (2016), Sharifi et al. (2016), Kennedy (2018), Tajika et al. (2018), El Qot (2018), Robert et al. (2018).

Subtaxa

Engonoceratidae, Forbesiceridae, Hoplitidae, Placenticeratidae (syn. Hypengonoceratidae), Schloenbachiidae

Synonymy list

Year	Name and author
1890	Hoplitaceaea Douvillé
1952	Hoplitaceae Wright
1970	Hoplitaceae Nagy p. 31
1975	Hoplitaceae Förster p. 210
1978	Hoplitaceae Cooper p. 73
1978	Hoplitaceae Wiedmann p. 666
1979	Hoplitaceae Cooper and Kennedy p. 235
1979	Hoplitaceae Summesberger p. 145
1981	Hoplitaceae Kennedy et al. p. 229
1984	Hoplitaceae Kennedy p. 38
1989	Hoplitaceae Klinger and Kennedy p. 266
1991	Hoplitaceae Kennedy and Simmons p. 132
1992	Hoplitaceaea Kennedy et al. p. 270
1995	Hoplitaceae Kennedy p. 410
1996	Hoplitaceae Wright et al. p. 114
1997	Hoplitaceae Cecca p. 327
1997	Hoplitaceae Kennedy et al. p. 4
1998	Hoplitaceae Kennedy et al. p. 7
1999	Hoplitaceae Kennedy and Summesberger p. 25
2005	Hoplitoidea Kennedy p. 357
2007	Hoplitoidea Kennedy and Latil p. 461
2007	Hoplitaceae Szives
2008	Hoplitoidea Aly et al. p. 46
2008	Hoplitoidea Kennedy et al. p. 38
2008	Hoplitoidea Kennedy et al. p. 123
2014	Hoplitoidea Mosavinia et al. p. 74
2016	Hoplitoidea Kennedy and Gale p. 269
2016	Hoplitaceae Sharifi et al. p. 597
2016	Hoplitoidea Shigeta p. 343
2018	Hoplitoidea El Qot p. 156
2018	Hoplitoidea Kennedy p. 678
2018	Hoplitaceae Robert et al. p. 9
2018	Hoplitoidea Tajika et al. p. 69

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
phylum	Mollusca

Rank	Name	Author
class	Cephalopoda	Cuvier 1797
subclass	Coleoidea	Bather 1888
order	Ammonoida	(Haeckel 1866)
order	Ammonitida	Haeckel 1866
suborder	Ammonitina	Hyatt 1889
superfamily	Hoplitoidea	(Douvillé 1890)
superfamily	Hoplitoidea	(Douvillé 1890)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. †Hoplitoidea Douvillé 1890

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Fm. †Engonoceratidae Hyatt 1900

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G. †Engonoceras Neumayr and Uhlig 1881

†Engonoceras belviderense Cragin 1894

†Buchiceras (Sphenodiscus) belviderensis serpentinus Cragin 1900

Invalid names: Engonoceras uddeni Cragin 1900 [synonym]

†Engonoceras complicatum Hyatt 1903

†Engonoceras duboisi Latil 1989

†Engonoceras elegans Kennedy et al. 1998

†Engonoceras emarginatum Cragin 1893

†Engonoceras gibbosum Hyatt 1903

†Engonoceras hilli Böhm 1898

†Engonoceras iris Spath 1924

†Engonoceras pierdenale Von Buch 1848

†Engonoceras retardum Hyatt 1903

†Engonoceras serpentinum Cragin 1900

†Engonoceras stolleyi Böhm 1898

†Engonoceras subjectum Hyatt 1903

†Engonoceras wilkinsoni Packard 1956

G. †Metengonoceras Hyatt 1903

†Metengonoceras acutum Hyatt 1903

†Metengonoceras ambiguum Hyatt 1903

†Metengonoceras aspenanum Reeside and Weymouth 1931

†Metengonoceras bedoti Grossouvre 1912

†Metengonoceras bravoense Böse 1928

†Metengonoceras dumbli Cragin 1893

Invalid names: Metengonoceras arnaudi Grossouvre 1912 [synonym], Metengonoceras douvillei Grossouvre 1912 [synonym], Metengonoceras nigeriense Furon 1935 [synonym], Metengonoceras tolveiense Grossouvre 1912 [synonym]

†Metengonoceras inscriptum Hyatt 1903

†Metengonoceras oliveirai Brito 1967

†Metengonoceras teigenense Cobban and Kennedy 1989

Invalid names: Epengonoceras Spath 1924 [synonym]

G. †Neolobites Fisher 1882

†Neolobites fourteaui Pervinquière 1907

Invalid names: Neolobites medeninensis Basse 1954 [synonym]

†Neolobites kummeli Benavides-Caceres 1956

†Neolobites peroni Hyatt 1903

†Neolobites vibrayeanus d'Orbigny 1841

Invalid names: Neolobites brancai Eck 1908 [synonym], Neolobites bussoni Collignon 1965 [synonym], Neolobites choffati Hyatt 1903 [synonym], Neolobites schweinfurthi Eck 1914 [synonym]

G. †Protengonoceras Hyatt 1900

†Protengonoceras gabbi Böhm 1898

†Protengonoceras neolobitoides Robert et al. 2018

†Protengonoceras planum Hyatt 1903

Fm. †Hoplitidae Douvillé 1890

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Subfm. †Anahoplitinae Breistroffer 1947

G. †Anahoplites Hyatt 1900

†Anahoplites gracilis Spath 1928

†Anahoplites picteti Spath 1925

†Anahoplites planus Mantell 1822

Invalid names: Ammonites splendens Sowerby 1815 [replaced]

†Anahoplites williamsi Maury 1936

†Anahoplites yakounensis Whiteaves 1900

G. †Anahoplitoides Casey 1961

G. †Lepthoplites

†Lepthoplites cantabrigiensis Spath 1926

†Lepthoplites gracilis Spath 1928

Invalid names: Semenovitinae Mirzoev 1967 [synonym]

Subfm. †Gastroplitinae Wright 1952

G. †Alopecoceras Kennedy 1978

G. †Anagastroplites Jeletzky 1980

†Anagastroplites tozeri Jeletzky 1980

G. †Gastroplites

†Gastroplites canadensis Whiteaves 1893

†Gastroplites subquadratus Nagy 1970

G. †Neogastroplites

†Neogastroplites americanus Reeside and Weymouth 1934

†Neogastroplites cornutus Whiteaves 1885

Invalid names: Kanabiceras wyomingense Reeside and Weymouth 1931 [synonym]

†Neogastroplites kamchatkensis Alabushev and Wiedmann 1994

G. †Paragastroplites Imlay 1961

G. †Pseudogastroplites Jeletzky 1980

†Pseudogastroplites arcticus Jeletzky 1980

†Pseudogastroplites nanus Jeletzky 1980

G. †Pseudopulchellia Imlay 1961

†Pseudopulchellia balkwilli Jeletzky 1980

†Pseudopulchellia flexicostata Imlay 1961

†Pseudopulchellia pattoni Imlay 1961

†Pseudopulchellia susukii Murphy and Rodda 1959

G. †Sokolovites Casey 1966

G. †Stelckiceras Jeletzky 1980

†Stelckiceras liardense Whiteaves 1889

G. †Stotticeras Jeletzky 1980

†Stotticeras crowense Jeletzky 1980

Subfm. †Hoplitinae Douvillé 1890

G. †Arrhaphoceras Whitehouse 1927

Subg. †Arrhaphoceras (Arrhaphoceras) Whitehouse 1927

†Arrhaphoceras (Arrhaphoceras) subtetragonum Spath 1928

Subg. †Arrhaphoceras (Praeschloenbachia) Scholz 1973

G. †Callihoplites

†Callihoplites auritus Sowerby 1816

†Callihoplites dorsetensis Spath 1928

†Callihoplites tetragonus Seeley 1865

Invalid names: Ammonites acanthonotus Seeley 1865 [synonym], Callihoplites atavus Spath 1928 [synonym], Callihoplites fuscus Spath 1928 [synonym], Callihoplites gymnus Spath 1928 [synonym], Callihoplites paradoxus Spath 1928 [synonym], Callihoplites pulcher Spath 1928 [synonym], Callihoplites senilis Spath 1928 [synonym]

†Callihoplites variabilis Spath 1927

G. †Dimorphoplites

†Dimorphoplites tethydis Bayle 1878

G. †Discohoplites

†Discohoplites coelonotus Seeley 1865

†Discohoplites simplex Wright and Wright 1949

†Discohoplites subfalcatus Semenov 1899

†Discohoplites valbonnensis Hébert and Munier-Chalmas 1875

G. †Epihoplites

Subg. †Epihoplites (Metaclavites) Casey 1965

†Epihoplites (Metaclavites) iphitus Spath 1926

Invalid names: Epihoplites trapezoidalis Luppov 1961 [synonym]

Invalid names: Gazdaganites [synonym]

G. †Euhoplites

†Euhoplites reesidei Maury 1936

G. †Hoplites

Subg. †Hoplites (Hoplites) Neumayr 1875

†Hoplites (Hoplites) obtusus Spath 1925

†Hoplites (Hoplites) svalbardensis Nagy 1970

Subg. †Hoplites (Isohoplites) Casey 1954

†Hoplites (Isohoplites) eodentatus Casey 1961

†Isohoplites steinmanni Jacob 1907

†Hoplites aguilearae Böse 1923

†Hoplites mirabiliformis Spath 1925

†Hoplites moutonianus d'Orbigny 1850

†Hoplites obtusenodosa Retowski 1893

†Hoplites symonensis Böse 1923

†Neoptychites (Hoplites) wohltmanni von Koenen 1897

G. †Hyphoplites Spath 1922

†Hyphoplites campichei Spath 1925

†Hyphoplites costosus Wright and Wright 1949

†Hyphoplites curvatus Mantell 1822

†Hyphoplites curvatus arausionensis Hébert and Munier-Chalmas 1875

†Hyphoplites falcatus Mantell 1822

†Hyphoplites falcatus aurora Wright and Wright 1949

G. †Pleurohoplites

†Pleurohoplites renauxianus d'Orbigny 1840

G. †Semenoviceras Wright et al. 1996

Subg. †Semenoviceras (Planihoplites) Saveliev 1992

†Semenoviceras (Planihoplites) mangyshlakense Saveliev 1960

†Semenoviceras (Planihoplites) pseudocoelonodum Semenov 1899

†Semenoviceras (Planihoplites) uhligi Semenov 1899

Subg. †Semenoviceras (Semenoviceras) Wright 1996

†Semenoviceras (Semenoviceras) baisunense Luppov 1961

†Semenoviceras (Semenoviceras) litschkovi Saveliev 1960

†Semenoviceras michalskii Semenov 1899

Invalid names: Hoplites pseudoauritus Semenov 1899 [synonym], Semenovites (Semenovites) tamalakensis Saveliev 1992 [synonym]

†Semenoviceras solidus Savelev 1960

Invalid names: Semenovites [replaced]

Subfm. †Leymeriellinae Wright 1952

G. †Epileymeriella

Subfm. †Sonneratiinae Destombes 1973

G. †Bucaillella Destombes 1973

G. †Eosonneratia Saveliev 1973

G. †Farnhamia Casey 1954

†Farnhamia farnhamensis Casey 1954

G. †Otohoplites Steinmann 1925

†Otohoplites auritiformes Spath 1925

†Otohoplites glyphus Casey 1965

†Otohoplites guersanti d'Orbigny 1841

†Otohoplites raulinianus d'Orbigny 1841

G. †Protohoplites Spath 1923

Subg. †Protohoplites (Hemisonneratia) Casey 1952

†Protohoplites (Hemisonneratia) puzosianus d'Orbigny 1841

Subg. †Protohoplites (Protohoplites) Spath 1923

†Protohoplites (Protohoplites) archiacianus d'Orbigny 1841

G. †Pseudosonneratia Spath 1925

G. †Sonneratia Bayle 1878

†Sonneratia crossi Anderson 1938

†Sonneratia dutempleana d'Orbigny 1850

†Sonneratia kitchini Spath 1925

†Sonneratia perrinsmithi Anderson 1938

†Sonneratia rotator Casey 1965

G. †Tetrahoplites

G. †Sulcohoplites Michailova and Saveliev 1994

Fm. †Placenticeratidae Hyatt 1900

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G. †Gissarites Iljin 1958

†Gissarites iljini Khakimov 1976

†Gissarites kysylchense Iljin 1958

G. †Hengestites

G. †Hoplitoplacenticeras Paulcke 1906

Subg. †Hoplitoplacenticeras (Hoplitoplacenticeras) Paulcke 1907

†Hoplites plasticus semicostatus Paulcke 1907

†Hoplitoplacenticeras (Hoplitoplacenticeras) coesfeldiense Schluter 1867

†Hoplitoplacenticeras (Hoplitoplacenticeras) howarthi Collignon 1970

Invalid names: Hoplitoplacenticeras antokazoense Collignon 1970 [synonym], Hoplitoplacenticeras besairei Collignon 1970 [synonym], Hoplitoplacenticeras trangahyense Collignon 1970 [synonym]

†Hoplitoplacenticeras (Hoplitoplacenticeras) marroti Coquand 1859

†Hoplitoplacenticeras (Hoplitoplacenticeras) plasticum Paulcke 1907

†Hoplites plasticus costatus Paulcke 1907

†Hoplites plasticus hauthali Paulcke 1907

†Hoplites plasticus laevis Paulcke 1907

†Hoplitoplacenticeras (Hoplitoplacenticeras) preyi Kennedy and Summesberger 1999

Subg. †Hoplitoplacenticeras (Lemfoerdiceras) Kennedy 1986

†Hoplitoplacenticeras (Lemfoerdiceras) lafresnayanus d'Orbigny 1841

†Hoplitoplacenticeras (Lemfoerdiceras) lemfoerdensis Schluter 1872

†Hoplitoplacenticeras costulosum Schlüter 1867

†Hoplitoplacenticeras dolbergense Schluter 1876

†Hoplitoplacenticeras fugen Matsumoto 1984

†Hoplitoplacenticeras monju Matsumoto 1982

†Hoplitoplacenticeras plasticum Paulcke 1906

†Hoplitoplacenticeras rarecostatum Khakimov 1976

†Hoplitoplacenticeras rejaudryi de Grossouvre 1894

Invalid names: Sonneratia rara de Grossouvre 1894 [synonym]

†Hoplitoplacenticeras vari Schlüter 1872

G. †Karamaites Sokolov 1965

†Karamaites grossouvrei Semenov 1899

G. †Metaplacenticeras Spath 1926

†Metaplacenticeras bowersi Anderson 1958

†Metaplacenticeras californicum Anderson 1902

†Metaplacenticeras pacificum Smith 1900

†Metaplacenticeras sanctaemonicae Waring 1917

†Metaplacenticeras subtilistriatum Jimbo 1894

G. †Placenticeras Meek 1876

†Placenticeras benningi Stephenson 1956

†Placenticeras bidorsatum Roemer 1841

Invalid names: Ammonites dujardin Schluter 1867 [synonym], Diplacomoceras canaliculatum Hyatt 1903 [synonym]

†Placenticeras cumminsi Cragin 1893

†Placenticeras fritschi de Grossouvre 1894

†Placenticeras intercalare Meek and Hayden 1860

†Placenticeras kaffrarium Etheridge 1904

Invalid names: Placenticeras umkwelanensis Etheridge 1904 [synonym]

†Placenticeras kolbajense Sokolov 1967

†Placenticeras maherndli Summesberger 1979

†Placenticeras mediasiaticum Luppov 1963

Invalid names: Turkmenites gissarensis Ilyin 1975 [synonym]

†Placenticeras memoriaschloenbachi Laube and Bruder 1887

†Placenticeras orbignyanum Geinitz 1849

†Placenticeras paraplanum Wiedmann 1978

†Placenticeras placenta DeKay 1828

†Placenticeras polyopsis Dujardin 1837

†Placenticeras pseudoplacenta Hyatt 1903

†Placenticeras semiornatum d'Orbigny 1850

†Placenticeras syrtale Morton 1834

Invalid names: Ammonites guadalupae Roemer 1849 [synonym], Placenticeras intermedium Johnson 1903 [synonym], Placenticeras newberryi Hyatt 1903 [synonym], Placenticeras planum Hyatt 1903 [synonym], Placenticeras rotundatum Johnson 1903 [synonym], Placenticeras sancarlosense Hyatt 1903 [synonym], Stantonoceras pseudocostatum Johnson et al. 1964 [synonym]

†Placenticeras tamulicum Blanford 1862

†Placenticeras vancouverense Meek 1861

†Placenticeras vredenburgi Sarkar 1966

†Placenticeras whitfieldi Hyatt 1903

Invalid names: Diplacomoceras Hyatt 1900 [synonym], Karamaiceras [synonym], Pseudoplacenticeras [synonym]

G. †Proplacenticeras

†Proplacenticeras stantoni Hyatt 1903

Invalid names: Placenticeras merenskyi Haughton 1930 [synonym]

G. †Stantonoceras

†Stantonoceras depressum Hyatt 1903

Invalid names: Baghiceratinae Chiplonkar and Ghare 1976 [invalid subgroup], Hypengonoceratidae Chiplonkar and Ghare 1976 [synonym], Rapidoplacenticeratinae Alabushev and Alabusheva 1988 [empty]

Fm. †Schloenbachiidae Parona and Bonarelli 1897

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G. †Schloenbachia Neumayr 1875

†Schloenbachia blanfordianus Stoliczka 1863

†Schloenbachia lenzi Szajnocha 1885

†Schloenbachia varians Sowerby 1817

†Schloenbachia varians subtuberculata Gerhardt 1897

Invalid names: Saltericeras [synonym]

Invalid names: Forbesiceratinae Wright 1952 [empty], Schloenbachiinae Parona and Bonarelli 1897 [empty]

Invalid names: Forbesiceridae Wright 1952 [empty]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

aragoniteo

Environment:

marinec

Locomotion:

fast-movingo

Life habit:

nektonico

Diet:

carnivoreo

Vision:

well-developedc

Created:

2005-01-28 02:27:03

Modified:

2009-10-08 14:14:55

Source: o = order, c = class

References: Kiessling 2004, Kiessling 2003

Age range: base of the Tithonian to the top of the Early/Lower Maastrichtian or 149.20000 to 66.00000 Ma

Collections (736 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Late/Upper Jurassic	161.5 - 143.1	Argentina (Mendoza)	Hoplites vetustus (137144)
Tithonian - Berriasian	149.2 - 137.05	Russian Federation	Delphinella obtusenodosa (213004)
Berriasian	143.1 - 137.05	Ukraine	Delphinella obtusenodosa (163877 163882)
Berriasian - Turonian	143.1 - 89.8	USA (California)	Sonneratia crossi (216097)
Valanginian	137.05 - 132.6	Mexico (Durango)	Hoplites sp. (87612)
Early/Lower Aptian	121.4 - 119.57	Japan	Hoplitaceae indet. (65019)
Aptian	121.4 - 113.2	Spain (Catalonia)	Hoplites desayesi (111535)
Aptian	121.4 - 113.2	United Kingdom (England)	Sonneratia kitchini (1823)
Aptian	121.4 - 113.2	Svalbard and Jan Mayen	Hoplites sp. (211839 212052)
Late/Upper Aptian	119.57 - 113.2	USA (Texas)	Metengonoceras inscriptum (84441)
Youngest occurrences
Time interval	Ma	Country or state	PBDB collection number
Middle Campanian	83.6 - 72.2	USA (South Dakota)	Placenticeras sp. (235333)
Middle Campanian	83.6 - 72.2	USA (Texas)	Placenticeras meeki, Placenticeras sp. (7467) Placenticeras meeki, Placenticeras sp., Placenticeras syrtale (7470)
Middle Campanian	83.6 - 72.2	USA (California)	Metaplacenticeras californicum (221105 221106 221107)
Middle Campanian	83.6 - 72.2	Japan	Metaplacenticeras subtilistriatum (181920 181921 181922)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	USA (New Jersey)	Placenticeras placenta (3870)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	USA (California)	Metaplacenticeras pacificum (215264)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	USA (New Jersey)	Placenticeras minor (221052) Placenticeras syrtale (57916)
Late/Upper Maastrichtian	72.2 - 66.0	USA (North Carolina)	Placenticeras sp. (7616)
Late/Upper Maastrichtian	72.2 - 66.0	France	Hoplitoplacenticeras lasfresnayanum (83648)
Maastrichtian	72.2 - 66.0	USA (Texas)	Placenticeras interealare (7549) Placenticeras meeki (7483 7531 7536 7548 7569)