PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Macroelongatoolithus carlylei

Reptilia - Theropoda - Elongatoolithidae

Taxonomy

Oolithes carlylensis was named by Jensen (1970). It is considered to be a form taxon. Its type specimen is BYU-E 200, an egg, and it is a 3D body fossil. Its type locality is Castle Dale, Jensen Locality 1, which is in a Cenomanian terrestrial shale in the Cedar Mountain Formation of Utah. It is the type species of Macroelongatoolithus.

It was recombined as Boletuoolithus carlylensis by Bray (1998); it was recombined as Macroelongatoolithus carlylei by Zelenitsky et al. (2000), Jin et al. (2007), Bonde et al. (2008), Kim et al. (2011), Tanaka et al. (2011), Huh et al. (2014), Tanaka et al. (2016), Krumenacker et al. (2017) and Simon et al. (2019); it was misspelled as Macroelongatoolithus carleylei by Krumenacker et al. (2022).

Synonyms

Longiteresoolithus xixiaensis was named by Wang and Zhou (1995). It is considered to be a form taxon. It is a 3D body fossil. Its type locality is Liuying, Yangcheng, which is in a Cenomanian/Turonian terrestrial siltstone in the Gaogou Formation of China.

It was recombined as Macroelongatoolithus xixiaensis by Jin et al. (2007), Jin (2008), Wang et al. (2010), Kim et al. (2011), Wang et al. (2012) and Huh et al. (2014); it was misspelled as Macroelongatoolithus xixia by Cousin (2002), Zhou and Feng (2002) and Liang et al. (2009); it was synonymized subjectively with Macroelongatoolithus carlylei by Zelenitsky et al. (2000) and Simon et al. (2019).

Synonymy list

Year	Name and author
1970	Oolithes carlylensis Jensen pp. 62-63 figs. 5, Pl. 1:1-2
1995	Longiteresoolithus xixiaensis Wang and Zhou p. 262
1998	Boletuoolithus carlylensis Bray p. 221
2000	Macroelongatoolithus carlylei Zelenitsky et al. p. 131–132
2002	Macroelongatoolithus xixia Cousin p. 44
2002	Longiteresoolithus xixiaensis Zhou and Feng p. 69
2007	Macroelongatoolithus carlylei Jin et al. p. 82
2007	Macroelongatoolithus xixiaensis Jin et al. p. 82
2008	Macroelongatoolithus carlylei Bonde et al. p. 255
2008	Macroelongatoolithus xixiaensis Jin p. 53
2009	Longiteresoolithus xixiaensis Liang et al. p. 1591
2010	Macroelongatoolithus xixiaensis Wang et al. p. 96
2011	Macroelongatoolithus carlylei Kim et al. p. 18
2011	Macroelongatoolithus xixiaensis Kim et al. p. 18
2011	Macroelongatoolithus carlylei Tanaka et al. p. 45
2012	Macroelongatoolithus xixiaensis Wang et al. p. 403
2014	Macroelongatoolithus carlylei Huh et al. p. 220
2014	Macroelongatoolithus xixiaensis Huh et al. pp. 220-221
2016	Macroelongatoolithus carlylei Tanaka et al. p. 353
2017	Macroelongatoolithus carlylei Krumenacker et al. p. 170
2019	Macroelongatoolithus carlylei Simon et al. p. 7–8
2022	Macroelongatoolithus carleylei Krumenacker et al. p. 1339

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida

Rank	Name	Author
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
order	Theropoda
family	Elongatoolithidae
genus	Macroelongatoolithus	Li et al. 1995
species	carlylei	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Macroelongatoolithus carlylei Jensen 1970

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Invalid names: Longiteresoolithus xixiaensis Wang and Zhou 1995 [synonym]

Diagnosis

Reference	Diagnosis
D. K. Zelenitsky et al. 2000	Same as for oogenus.
M. Huh et al. 2014 (Macroelongatoolithus xixiaensis)	Large, elongate eggs 39 – 52 cm long and 13 – 18 cm wide. Elongation index ranges from 2.6 to 3.9. Continuous layer to mammillary layer ratio variable, with a distinct, undulating boundary between layers. Eggshell thickness ranges from 1.5 to 4.45 mm including surface ornamentation. Shell surface covered with coarse lineartuberculate ornamentation at equator grading to dispersituberculate at the poles. Eggs arranged in pairs within single-layered ring-shaped clutch with a central space devoid of eggs (modified after Jin et al. 2007).
D. J. Simon et al. 2019	Same as for oogenus.

Measurements

No measurements are available

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

solitaryo

Environment:

terrestrialo

Locomotion:

actively mobileo

Life habit:

ground dwellingo

Diet:

carnivoreo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2004-12-14 12:50:08

Modified:

2004-12-14 14:50:08

Source: o = order

Reference: Marsh 1875

Age range: base of the Late/Upper Albian to the top of the Campanian or 105.30000 to 72.10000 Ma

Collections (13 total)

Time interval	Ma	Country or state	Original ID and collection number
Albian	113.0 - 100.5	China (Zhejiang)	Macroelongatoolithus xixiaensis (86428)
Late/Upper Albian - Early/Lower Cenomanian	105.3 - 93.5	USA (Utah)	Macroelongatoolithus carlylei (204699)
Late/Upper Albian - Early/Lower Cenomanian	105.3 - 93.5	USA (Idaho)	Macroelongatoolithus carlylei (204701 204702)
Cenomanian	100.5 - 93.9	USA (Nevada)	Macroelongatoolithus carlylei (181585)
Cenomanian - Turonian	100.5 - 89.8	China (Henan)	Longiteresoolithus xixiaensis (178692 232202)
Early/Lower Cenomanian	99.6 - 93.5	USA (Utah)	Macroelongatoolithus carlylei (14399 117467 204698) Oolithes carlylensis (type locality: 52973)
Santonian - Campanian	86.3 - 72.1	China (Henan)	Longiteresoolithus xixiaensis (232203)
Campanian	83.6 - 72.1	South Korea (Jeollanam-do)	Macroelongatoolithus xixiaensis (232503)