PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Grallator limosus

Reptilia - Theropoda - Grallatoridae

Taxonomy

Grallator limosus was named by Zhen et al. (1986). It is considered to be a form taxon. Its type specimen is BPV-FP1, a footprint, and it is a trace fossil. Its type locality is Xiiyang, Jinning tracksites (BPV), which is in a Hettangian terrestrial horizon in the Fengjiahe Formation of China.

Sister species lacking formal opinion data

G. pisanus

Synonymy list

Year	Name and author
1986	Grallator limosus Zhen et al. p. 2 fig. 2–3
1989	Grallator limnosus Zhen et al. p. 187
2008	Grallator limnosus Fujita et al. p. 33
2009	Grallator limnosus Xing et al. p. 23
2013	Grallator limnosus Lockley et al. p. 6
2014	Grallator limnosus Xing et al. p. 344
2017	Grallator limnosus Xing et al. p. 115
2021	Grallator limnosus Li et al. p. 14

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida

Rank	Name	Author
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
order	Theropoda
family	Grallatoridae
genus	Grallator
species	limosus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Grallator limosus Zhen et al. 1986

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Diagnosis

Reference	Diagnosis
S. Zhen et al. 1986	The same as the characters of the genus (Lull 1904). The length of pes and pace is in a ratio of about 1:3.6. The step-angle is about 147°. The angles between phalanges are II-20°-III-15°-IV. The outline of the footprints is compact.
S. Zhen et al. 1989	Bipedal tridactyl tracks, tail trace absent. Claws accuminate, moderately developed. The lengths of the pes and pace are in a ratio of about 1:3.6. The step angle is 147°. The angles between the toes are II-20°-III-15°-IV. Length of footprints is 27.0–28.0 cm anf the width is 17.0–20.0 cm. The pace is 95.0–103.5 cm, and the stride is 190.0 cm.

Measurements

No measurements are available

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

solitaryo

Environment:

terrestrialo

Locomotion:

actively mobileo

Life habit:

ground dwellingo

Diet:

carnivoreo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2004-12-14 12:50:08

Modified:

2004-12-14 14:50:08

Source: o = order

Reference: Marsh 1875

Age range: Hettangian or 201.30000 to 199.30000 Ma

Collections: one only

Time interval	Ma	Country or state	Original ID and collection number
Hettangian	201.3 - 199.3	China (Yunnan)	Grallator limosus (type locality: 52848)