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Horaiclavidae
Taxonomy
Horaiclavidae was named by Bouchet et al. (2011). It is extant. Its type is Horaiclavus.
It was assigned to Conoidea by Bouchet et al. (2011).
It was assigned to Conoidea by Bouchet et al. (2011).
Subtaxa
Anacithara, Anguloclavus, Aoteadrillia, Austrocarina, Austrodrillia, Buchema, Carinapex, Ceritoturris, Coronacomitas, Cytharoclavus, Darrylia, Graciliclava, Haedropleura, Horaiclavus (type genus), Inkinga, Inodrillia (syn. Inodrillina, Inodrillara), Iwaoa, Marshallena (syn. Sugitanitoma), Mauidrillia, Micropleurotoma, Naskia, Nquma, Paradrillia (syn. Alticlavatula, Vexitomina), Pseudexomilus, Psittacodrillia, Striatoguraleus, Thelecythara
Synonymy list
| Year | Name and author |
|---|---|
| 2011 | Horaiclavidae Bouchet et al. pp. 293 - 296 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Horaiclavidae Bouchet et al. 2011
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G. Anacithara Hedley 1922
G. Anguloclavus Shuto 1983
G. Aoteadrillia Powell 1942
G. Austrocarina Laseron 1954
G. Buchema Corea 1934
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Buchema interstrigata Smith 1882
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Invalid names: Drillia (Neodrillia) euphanes Melvill 1923 [synonym]
G. Carinapex Dall 1924
G. †Ceritoturris Dall 1924
G. Coronacomitas Shuto 1983
G. Cytharoclavus Kuroda and Oyama in Kuroda Habe and Oyama 1971
G. Darrylia Garcia 2008
G. Graciliclava Shuto 1983
G. Haedropleura Monterosato 1883
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†Haedropleura orientalis Vredenburg 1923
†Haedropleura septangularis Montagu 1803
G. Horaiclavus Oyama 1954
G. Inkinga Kilburn 1988
G. Inodrillia Bartsch 1943
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†Inodrillia aepynota Dall 1889
Invalid names: Inodrillara Bartsch 1943 [synonym], Inodrillina Bartsch 1943 [synonym]
G. Iwaoa Kuroda 1940
G. Micropleurotoma Thiele 1929
G. Naskia Sysoev and Ivanov 1985
G. Nquma Kilburn 1988
G. Paradrillia Makiyama 1940
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Invalid names: Alticlavatula MacNeil 1960 [synonym], Vexitomina Powell 1942 [synonym]
G. Pseudexomilus Powell 1944
G. Psittacodrillia Kilburn 1988
G. Striatoguraleus Kilburn 1994
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| P. Bouchet et al. 2011 | Shell generally small, 5–25 mm, usually 7–15 mm high, shortly claviform, with relatively low spire and a short, truncated, poorly differentiated siphonal canal. Subsutural ramp usually poorly differentiated. Axial sculpture almost always present, usually as strong sinuate ribs. Spiral sculpture normally weak or obsolete, often with glossy shell surface. Anal sinus on subsutural slope, weak to moderately deep, often constrained by callus. Protoconch of up to 3.5 medially carinate but otherwise smooth whorls when multispiral, but usually paucispiral and smooth. Operculum with terminal nucleus. Radular formula: 1-0-0-0-1. Marginal teeth duplex, with lanceolate major limb and usually narrow accessory limb, which is inserted in a shallow socket. Major limb often with angulation lateral to the place of accessory limb insertion. Rarely (some Paradrillia, cf. Kilburn, 1988, Fig. 17; Inkinga, Fig. 19B) the additional limb is of similar size to the major limb and the teeth become trough-shaped in transverse section, with a collar near the base (arrows in Fig. 19B). In several species of Horaiclavus the radular apparatus is absent. |
Measurements
No measurements are available
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| Source: superf = superfamily, c = class | |||||
| References: Kiessling 2004, Hendy et al. 2009 | |||||
Age range
Maximum range based only on fossils: base of the Thanetian to the top of the Late/Upper Pleistocene or 59.24000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 56.0 Ma
Minimum age of oldest fossil (stem group age): 56.0 Ma
Collections (65 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Thanetian | India | Haedropleura orientalis (145510) | |
| Thanetian - Ypresian | Australia (Victoria) | Mauidrillia otwayensis (85124) | |
| Middle Eocene | Indonesia | Paradrillia boehmi (70082) | |
| Bortonian | New Zealand | Marshallena serotina (64733) | |
| Bartonian | Russian Federation (Kamchatka) | Marshallena tigilensis, Micropleurotoma sp. (5946) | |
| Bartonian - Priabonian | Australia (Western Australia) | Mauidrillia aldingensis (179825) | |
| Kaiatan | New Zealand | Marshallena esdailei (67839) | |
| Late/Upper Eocene | New Zealand (Canterbury) | Marshallena neozelanica (45037 45041) | |
| Early/Lower Oligocene | New Zealand | Aoteadrillia calliomorpha (67925) | |
| Rupelian | Germany | Mauidrillia sp. (95898) | |
| Oligocene | New Zealand | Marshallena sp. (45023) | |
| Late/Upper Oligocene | New Zealand | Marshallena anomala, Mauidrillia inaequalis (67864) Mauidrillia inaequalis (67861) | |
| Early/Lower Miocene | Germany (North Rhein-Westfalia) | Ceritoturris hannoverana (62882 62883) | |
| Early/Lower Miocene | New Zealand (Dannevirke) | Marshallena sp. (35647) | |
| Early/Lower Miocene | India | Paradrillia serana (72084) | |
| Early/Lower Miocene - Middle Miocene | Germany (North Rhein-Westfalia) | Ceritoturris hannoverana (62884) | |
| Miocene | Germany (North Rhein-Westfalia) | Ceritoturris hannoverana (62899) | |
| Otaian | New Zealand | Anacithara axialis (77488) | |
| Burdigalian | Italy | Haedropleura sp. 1 (178108) | |
| Burdigalian | USA (Delaware) | Inodrillia whitfieldi (18579) | |
| Altonian | New Zealand | Anacithara n. sp., Aoteadrillia callimorpha, Marshallena anomala, Marshallena sp., Vexitomina sp., Mauidrillia supralaevis, Mauidrillia clavicula, Mauidrillia n. sp. (67897) Mauidrillia praecophinoides, Mauidrillia costifer, Mauidrillia angustata (67900) Mauidrillia supralaevis, Mauidrillia praecophinoides (67898) Mauidrillia unilirata, Mauidrillia angustata (67899) | |
| Langhian | Italy | Haedropleura sp. 1 (178116) | |
| Langhian - Serravallian | Netherlands | Haedropleura maitreja (186883 186884) | |
| Middle Miocene | Indonesia | Paradrillia ermelingi (70085) | |
| Middle Miocene | Germany (North Rhein-Westfalia) | Ceritoturris hannoverana (61905 61906 62887) | |
| Waiauan | New Zealand | Mauidrillia occidentalis (63089 63090) | |
| Tortonian | Denmark | Haedropleura maitreja (98312) | |
| Tortonian | Panama (Darien) | Thelecythara sp. (136122) | |
| Late/Upper Miocene | Venezuela | Buchema sp. (92751) | |
| Late/Upper Miocene - Early/Lower Pliocene | Japan (Okinawa) | Mauidrillia kachabaruensis (42181) | |
| Early/Lower Pliocene | Japan (Okinawa) | Clavatula kakegawensis (42229) | |
| Zanclean | Italy | Haedropleura bucciniformis (71213) | |
| Pliocene | USA (Florida) | Buchema sp. (146875) | |
| Zanclean - Piacenzian | Italy | Haedropleura bucciniformis (72068) Haedropleura bucciniformis, Haedropleura septangularis (71103) | |
| Zanclean - Piacenzian | Italy (Siena) | Haedropleura secalina, Haedropleura septangularis (71102) | |
| Late/Upper Pliocene | Costa Rica (Limon) | Thelecythara sp. (79843) | |
| Late/Upper Pliocene | Jamaica | Thelecythara mucronata (58522 69090 69094) | |
| Piacenzian | Italy | Haedropleura bucciniformis (71212) Haedropleura bucciniformis, Haedropleura secalina (71130) Haedropleura secalina, Haedropleura septangularis (71101) Haedropleura secalina, Haedropleura sp. 1, Haedropleura septangularis (71154) | |
| Early/Lower Pleistocene | USA (Florida) | Thelecythara floridana, Cymatosyrinx aepynota (79331) | |
| Early/Lower Pleistocene | Fiji | Paradrillia kakegawaensis (94832) | |
| Pleistocene | Philippines (Panay) | Paradrillia inconstans (41651) | |
| Late/Upper Pleistocene | Japan | Paradrillia inconstans (97961) | |
| Late/Upper Pleistocene | Portugal | Haedropleura septangularis (122284 122286 122288 122290 122291 122292) | |
| Late/Upper Pleistocene | Portugal (Azores) | Haedropleura sp. (128470) |