PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Eubaeninae

Reptilia - Baenidae

Taxonomy

Eubaeninae was named by Williams (1950).

It was reranked as the unranked clade Eubaeninae by Joyce and Lyson (2015), Lyson et al. (2016).

It was assigned to Baenidae by Gaffney (1972), Archibald and Hutchison (1979); and to Baenodda by Joyce and Lyson (2015), Lyson et al. (2016).

Subtaxa

Baena, Baena marshi, Boremys, Chisternon, Chisternon interpositum, Denazinemys, Goleremys, Stygiochelys

Synonyms

Baena marshi was named by Hay (1904). Its type specimen is YPM 299, a partial shell (Hay mentioned but did not figure carapace elements, but the type currently consists only of the plastron), and it is a 3D body fossil. Its type locality is south side of Lightning Creek, which is in a Maastrichtian terrestrial horizon in the Lance Formation of Wyoming.

It was considered a nomen dubium by Lyson and Joyce (2010), Joyce and Lyson (2015).
Chisternon interpositum was named by Hay (1909). Its type specimen is USNM 6058, a partial shell (anterior portions of carapace and plastron), and it is a 3D body fossil. Its type locality is About 10 or 15 miles west of north of Red Lodge, which is in a Paleocene terrestrial horizon in the Fort Union Formation of Montana.

It was considered a nomen dubium by Gaffney (1972), Joyce and Lyson (2015).

Synonymy list

Year	Name and author
1904	Baena marshi Hay
1908	Baena marshi Hay p. 62 figs. Plate 8, fig. 2; text-fig. 39
1909	Chisternon interpositum Hay pp. 194-195
1930	Baena marshi Hay p. 72
1930	Chisternon interpositum Hay p. 73
1950	Eubaeninae Williams
1972	Eubaeninae Gaffney pp. 248, 259
1972	Baena marshi Gaffney p. 301
1979	Eubaeninae Archibald and Hutchison
2015	Eubaeninae Joyce and Lyson
2016	Eubaeninae Lyson et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha

Rank	Name	Author
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
—	Testudinata	(Oppel 1811)
—	Paracryptodira
family	Baenidae	Cope 1882
—	Baenodda	()
—	Eubaeninae	(Williams 1950)
—	Eubaeninae	(Williams 1950)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. †Eubaeninae Williams 1950

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G. †Baena Leidy 1870

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†Baena affinis Leidy 1871

Invalid names: Baena inflata Gilmore 1915 [synonym], Baena riparia Hay 1908 [synonym]

†Baena arenosa Leidy 1870

Invalid names: Baena clara Hay 1908 [synonym], Baena emiliae Hay 1908 [synonym], Baena sima Hay 1908 [synonym]

†Baena escavada Hay 1908

†Baena hayi Gilmore 1916

G. †Boremys Lambe 1906

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†Boremys grandis Gilmore 1935

†Boremys pulchra Lambe 1906

Invalid names: Boremys albertensis Gilmore 1919 [synonym]

G. †Chisternon Leidy 1872

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†Chisternon undatum Leidy 1871

Invalid names: Baena gigantea Gilmore 1915 [synonym], Baena hebraica Cope 1872 [synonym]

G. †Denazinemys Lucas and Sullivan 2006

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†Denazinemys nodosa Gilmore 1916

G. †Goleremys Hutchison 2004

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†Goleremys mckennai Hutchison 2004

G. †Stygiochelys Gaffney and Hiatt 1971

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†Stygiochelys estesi Gaffney and Hiatt 1971

Invalid names: Baena marshi Hay 1904 [nomen dubium], Chisternon interpositum Hay 1909 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

terrestrialuc

Locomotion:

actively mobilec

Created:

2017-04-17 15:15:35

Modified:

2017-04-17 15:15:35

Source: c = class, subp = subphylum, uc = unranked clade

References: Hendy et al. 2009, Kiessling 2004, Carroll 1988

Age range: base of the Judithian to the top of the Lutetian or 83.60000 to 41.03000 Ma

Collections (172 total)

Time interval	Ma	Country or state	Original ID and collection number
Campanian	83.6 - 72.2	USA (New Mexico)	Denazinemys nodosa (240794)
Campanian	83.6 - 72.2	Canada (Alberta)	Boremys albertensis (128460) Boremys pulchra (128459 177142)
Early/Lower Campanian	83.6 - 72.2	USA (North Carolina)	Baena sp. (106277)
Early/Lower Campanian	83.6 - 72.2	USA (Texas)	Denazinemys sp. (200394 200399 200400)
Early/Lower Campanian - Middle Campanian	83.6 - 72.2	USA (Utah)	Denazinemys nodosa (167916)
Judithian	83.6 - 72.2	USA (Texas)	Baena sp. (14489) Denazinemys sp. (52463 151416 222815)
Judithian	83.6 - 72.2	USA (Wyoming)	Boremys sp. (14458)
Late/Upper Campanian	83.6 - 72.2	USA (Montana)	Boremys pulchra (14452)
Late/Upper Campanian	83.6 - 72.2	Canada (Alberta)	Boremys sp. (98674)
Late/Upper Campanian	83.6 - 72.2	USA (New Mexico)	Baena nodosa (52052 52874 100675 128561 128562 128563 196539) Baena nodosa, Boremys grandis (76760) Baena sp. (196540) Boremys grandis (53981) Boremys sp. (154570) Denazinemys nodosa (47820 53899 62317 62318 76106 76763 96813 96814 138830 138835 144519 154366 154367 154371 154374 154382 154386 154395 154423 154433 154462 154571 154584 154585 154586 154587 154588 154589 154590 154591 154592 154593 154594 154595 154596 154597 154598 222078) Denazinemys nodosa, Baenidae indet. (76105) Denazinemys sp. (138832)
Late/Upper Campanian	83.6 - 72.2	Mexico (Coahuila)	Denazinemys nodosa (170939)
Late/Upper Campanian	83.6 - 72.2	USA (Utah)	Boremys grandis (187677 187697) Boremys grandis, Denazinemys nodosa (187674) Boremys sp. (219712) Denazinemys nodosa (187655 187660 187665 187678 187680 187683 187702 187706 187712 231039) Denazinemys sp. (192256)
Middle Campanian	83.6 - 72.2	USA (Utah)	Baena sp. (167520)
Middle Campanian	83.6 - 72.2	USA (Texas)	Denazinemys nodosa (240118)
Middle Campanian	83.6 - 72.2	USA (Montana)	Boremys sp. (14437)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	Canada (Saskatchewan)	Boremys sp. (14494)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	USA (Montana)	Boremys sp. (14434 14457 14462 14490 222122)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	USA (Texas)	Baena nodosa (212185)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	Canada (Alberta)	Boremys sp. (52249)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	USA (Texas)	Denazinemys nodosa (240119)
Middle Campanian - Early/Lower Maastrichtian	83.6 - 66.0	USA (Texas)	Denazinemys sp. (240127)
Lancian	72.2 - 66.0	USA (Montana)	Stygiochelys estesi (14654)
Lancian	72.2 - 66.0	USA (Wyoming)	Baena sp. (14519 14579) Stygiochelys sp. (14505)
Lancian	72.2 - 66.0	USA (North Dakota)	Boremys sp. (14610)
Late/Upper Maastrichtian	72.2 - 66.0	USA (North Dakota)	Boremys sp. (45099 45102 45118 45137)
Late/Upper Maastrichtian	72.2 - 66.0	USA (Wyoming)	Baena hayi (122902) Baena marshi (52683)
Late/Upper Maastrichtian	72.2 - 66.0	USA (Texas)	Denazinemys nodosa (240126) Denazinemys sp. (189806)
Late/Upper Maastrichtian	72.2 - 66.0	USA (Montana)	Boremys sp. (229083)
Late/Upper Maastrichtian	72.2 - 66.0	Canada (Saskatchewan)	Baena sp. 1, Baena sp. 2 (45675)
Maastrichtian	72.2 - 66.0	USA (Montana)	Stygiochelys estesi (123100)
Maastrichtian	72.2 - 66.0	USA (North Dakota)	Boremys sp. (45135)
Pu1	66.0 - 63.8	USA (North Dakota)	Boremys sp. (105890)
Puercan	66.0 - 63.8	USA (Colorado)	Boremys sp. (209889) Stygiochelys estesi (14678 168453) Stygiochelys sp. (206185)
Paleocene	66.0 - 56.0	USA (Montana)	Chisternon interpositum (128448)
Torrejonian	63.8 - 60.9	USA (New Mexico)	Baena escavada (14834) Goleremys mckennai (232086)
Torrejonian	63.8 - 60.9	USA (Wyoming)	Eubaeninae indet. (128435)
Tiffanian	60.9 - 56.2	USA (California)	Goleremys mckennai (139917)
Ypresian - Lutetian	56.0 - 41.03	USA (Wyoming)	Baena arenosa (193357)
Wasatchian	54.9 - 50.5	USA (Wyoming)	Baena arenosa (15661) Baena sp. (15793 59890) Chisternon sp. (59892)
Wasatchian	54.9 - 50.5	USA (New Mexico)	Baena arenosa (176881)
Bridgerian	50.5 - 46.2	USA (Wyoming)	Baena arenosa (16229 16253 16262 16265 16647 27269 70257) Baena arenosa, Baena affinis (16201) Baena arenosa, Baena sima, Baena riparia, Chisternon undatum, Chisternon hebraicum (16228) Baena arenosa, Baena undata (131861) Baena sima (16255) Baena undata (16211 128447) Chisternon sp. (74504) Chisternon undatum (16226 106325) Chisternon undatum, Baena hebraica (130809)
Bridgerian - Early/Lower Uintan	50.5 - 39.7	USA (Wyoming)	Baena arenosa (128443)
Middle Eocene	48.07 - 37.71	USA (Wyoming)	Baena arenosa, Chisternon undatum, Baena undata (81455)
Early/Lower Uintan	46.2 - 39.7	USA (Utah)	Baena arenosa (16572 109207 186888) Chisternon undatum (186887)
Late/Upper Uintan	46.2 - 39.7	Canada (Saskatchewan)	Baena sp. (16626)
Uintan	46.2 - 39.7	USA (Utah)	Baena arenosa (185685 186889 186891 186892 186893 186895 186897 222799 222805) Baena arenosa, Chisternon undatum (186890 186894 186896) Baena emiliae (16589 185687 185688 185689 185691) Baena emiliae, Baena inflata, Baena gigantea (16646) Baena inflata (185690)