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Betaneospirifer moosakhailensis
Taxonomy
Spirifera moosakhailensis was named by Davidson (1862). Its type specimen is BMNH 82371 and is a 3D body fossil. Its type locality is Moosakhail (Fleming coll), which is in a Wuchiapingian marine limestone in the Wargal Formation of Pakistan.
It was recombined as Spirifer moosakhailensis by Grabau (1931); it was recombined as Spirifer musakheylensis by Astre (1934); it was recombined as Spirifer (Neospirifer) musakheylensis by Reed (1944); it was recombined as Neospirifer moosakhailensis by Waterhouse (1966), Waterhouse (1978), Lee and Gu (1980) and Shi and Shen (1997); it was recombined as Neospirifer (Neospirifer) moosakhailensis by Shen et al. (2003); it was misspelled as Neospirifer musakheylensis by Shimizu (1981); it was recombined as Betaneospirifer moosakhailensis by Waterhouse (2004).
It was recombined as Spirifer moosakhailensis by Grabau (1931); it was recombined as Spirifer musakheylensis by Astre (1934); it was recombined as Spirifer (Neospirifer) musakheylensis by Reed (1944); it was recombined as Neospirifer moosakhailensis by Waterhouse (1966), Waterhouse (1978), Lee and Gu (1980) and Shi and Shen (1997); it was recombined as Neospirifer (Neospirifer) moosakhailensis by Shen et al. (2003); it was misspelled as Neospirifer musakheylensis by Shimizu (1981); it was recombined as Betaneospirifer moosakhailensis by Waterhouse (2004).
Synonymy list
| Year | Name and author |
|---|---|
| 1862 | Spirifera moosakhailensis Davidson p. 28 figs. Pl 2, fig 2 |
| 1883 | Spirifer musakheylensis Waagen p. 512 figs. Pl 45 |
| 1892 | Spirifer musakheylensis Rothpletz p. 79 figs. Pl 9, figs 1, 2 |
| 1897 | Spirifer musakhelensis Diener |
| 1903 | Spirifera musakheylensis Etheridge Jr. p. 12 |
| 1913 | Spirifer musakheylensis Mansuy p. 57 figs. Pl 5, fig 6 |
| 1931 | Spirifer moosakhailensis Grabau p. 168 figs. Pl 23, figs 5-8 |
| 1934 | Spirifer musakheylensis Astre p. 74 figs. Pl 8, figs 17,24 |
| 1944 | Spirifer (Neospirifer) musakheylensis Reed p. 196 |
| 1966 | Neospirifer moosakhailensis Waterhouse p. 34 figs. Pl 8, figs 1,2; pl 9, figs 1,4; pl 10, figs 1,2 |
| 1978 | Neospirifer moosakhailensis Waterhouse p. 57 figs. Pl 6, figs 19,20 |
| 1980 | Neospirifer moosakhailensis Lee and Gu p. 412 figs. Pl 177, figs 7,10 |
| 1981 | Neospirifer musakheylensis Shimizu p. 80 figs. Pl 8, fig 24 |
| 1997 | Neospirifer moosakhailensis Shi and Shen p. 51 figs. 7B-D |
| 2003 | Neospirifer (Neospirifer) moosakhailensis Shen et al. figs. 6.1-6.5 |
| 2004 | Betaneospirifer moosakhailensis Waterhouse |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Betaneospirifer moosakhailensis Davidson 1862
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Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| J. B. Waterhouse 1966 | Transverse Neospirifer with moderately low ventral umbo and moderately high ventral interarea, alate, with rounded anterior-lateral extremities, sulcus shallow and V-shaped in profile posteriorly, wide and more U-shaped anteriorly, fold moderately high and narrow, plicae well developed, costae fine to strong, moderately differentiated, crests rounded, concentric lamellae well developed. |
Measurements
| part | N | mean | minimum | maximum | median | s.d. | |
|---|---|---|---|---|---|---|---|
| shell height | 3 | 23.6 | 14.0 | 37.4 | 25.0 | 11.7 |
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| Source: o = order, c = class, p = phylum, uc = unranked clade | |||||
| References: Aberhan et al. 2004, Nesnidal et al. 2013 | |||||
Age range: base of the Sakmarian to the top of the Changhsingian or 293.52000 to 252.17000 Ma
Collections (143 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Asselian - Sakmarian | Laos | Spirifer musakheylensis (233571) | |
| Sakmarian | Australia (Western Australia) | Spirifera musakheylensis (101221) | |
| Artinskian | Australia (Western Australia) | Spirifer musakheylensis (131883) | |
| Kungurian | Australia (Western Australia) | Spirifera musakheylensis (131881) | |
| Roadian - Wordian | Vietnam | Spirifer musakheylensis (139251) | |
| Roadian - Wordian | Myanmar | Spirifer fasciger (208999) | |
| Wordian | Mongolia | Neospirifer moosakhailensis (109034) | |
| Wordian | China (Jilin) | Neospirifer moosakhailensis (111179) | |
| Wordian | Pakistan | Spirifer musakheylensis (124354) | |
| Wordian | Canada (British Columbia) | Spirifer moosakhailensis (62320) | |
| Wordian - Capitanian | China (Heilongjiang) | Neospirifer moosakhailensis (111166) | |
| Capitanian | Pakistan | Spirifer musakheylensis (119552 125883 127032 127033) | |
| Wuchiapingian | India | Spirifer musakheylensis (228446 230614 230621 230627 230633) | |
| Wuchiapingian | Pakistan | Spirifer musakheylensis (88933 89058 90250 90253 119527 119537 119541 119550 119851 124292 124357 124358 125887 125891 125892 125895 125896 125897 125898 125900 125901 125902 125903 125907 125908 125909 125910 125912 125913 125918 125923 125961 125963 126002 126004 126005 126006 126007 126008 126011 126263 126264 126265 126267 126268 126269 126270 126272 126278 126280 126281 126293 126294 126295 126296 126299 126302 126341 126342 126343 126345 126452 126453 126458 126459 126460 126463 126691 126692 126693 126738 126881 126892 126893 126937 126938 126984 126993 126995 127004 127005 127006 127007 127016 127025 127026 127027 127028 127030) Spirifera moosakhailensis (type locality: 117776 117779) | |
| Wuchiapingian | Indonesia | Spirifer musakheylensis (132787) | |
| Wuchiapingian | India (Kashmir) | Neospirifer musakheylensis (82586) | |
| Wuchiapingian | Madagascar (Diana) | Spirifer musakheylensis (81987) | |
| Lopingian | India | Spirifer moosakhailensis (230849 230853) Spirifer musakheylensis (230330) | |
| Wuchiapingian - Changhsingian | Pakistan | Spirifer musakheylensis (126704 126705 126707 126740 126743 126746 126747 126751 126762 126998 127017 127023) Spirifera moosakhailensis (117777) | |
| Changhsingian | Nepal | Neospirifer moosakhailensis (43152 43154 43157 43158 43159 120444 120445 120446 120448 120451) | |
| Changhsingian | Pakistan | Spirifer musakheylensis (89064 119608 126934) |