PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Hirtoscala tincta (tinted wentletrap)

Gastropoda - Epitoniidae

Taxonomy

Scalaria tincta was named by Carpenter (1864). It is extant.

It was recombined as Scala tincta by Arnold (1903); it was recombined as Epitonium (Nitidiscala) tinctum by Durham (1937); it was recombined as Epitonium tinctum by Woodring and Bramlette (1950); it was recombined as Nitidiscala tincta by DuShane (1979), Squires et al. (2006); it was recombined as Hirtoscala tincta by Hendy (2021).

Sister species lacking formal opinion data

H. mitraeforme, H. replicata

Synonyms

Epitonium (Nitidiscala) eelense was named by Durham (1937). Its type specimen is UCMP 30147 and is a 3D body fossil.

It was synonymized subjectively with Nitidiscala tincta by DuShane (1979).
Epitonium (Nitidoscala) continuatum was named by Oldroyd (1925) [In occurrence list as "Epitonium (Nitidoscala) [sic] continuatum" but in species description and figure captions as "Epitonium continuatum"]. Its type specimen is USNM 352383 and is a 3D body fossil. Its type locality is Nob Hill cut, San Pedro, which is in a Pleistocene marine horizon in the San Pedro Sand Formation of California.

It was recombined as Epitonium (Nitidiscala) continuatum by Oldroyd (1925); it was synonymized subjectively with Epitonium tinctum by Hendy (2021).
Scalaria subcoronata was named by Carpenter (1864). It is extant.

It was recombined as Epitonium (Nitidoscala) subcoronatum by Olrdoyd (1925); it was synonymized subjectively with Nitidiscala tincta by DuShane (1979); it was synonymized subjectively with Epitonium tinctum by Hendy (2021).
Epitonium continuatum was named by Oldroyd (1925). It is extant.

It was synonymized subjectively with Nitidiscala tincta by DuShane (1979).
Epitonium (Nitidiscala) tinctum bormanni was named by Strong (1941). It is extant.

It was synonymized subjectively with Nitidiscala tincta by DuShane (1979).

Synonymy list

Year	Name and author
1864	Scalaria subcoronata Carpenter p. 660
1864	Scalaria tincta Carpenter p. 660
1903	Scala tincta Arnold
1925	Epitonium continuatum Oldroyd pp. 13, 35 figs. plt. 2,fig. 10.
1925	Epitonium (Nitidiscala) continuatum Oldroyd pp. 35 - 36 figs. Plate 2, fig. 10; Plate 3, fig. 5
1925	Epitonium (Nitidoscala) continuatum Oldroyd pp. 35 - 36 figs. Plate 2, fig. 10; Plate 3, fig. 5
1925	Epitonium (Nitidoscala) subcoronatum Olrdoyd
1937	Epitonium (Nitidiscala) eelense Durham p. 488 figs. Plate 56, figure 7
1937	Epitonium (Nitidiscala) tinctum Durham p. 488
1941	Epitonium (Nitidiscala) tinctum bormanni Strong p. 47
1950	Epitonium tinctum Woodring and Bramlette p. 48
1979	Nitidiscala tincta DuShane p. 117 figs. Figures 41, 45, 46, 47, 48
2006	Nitidiscala tincta Squires et al. p. 6
2021	Hirtoscala tincta Hendy

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
phylum	Mollusca

Rank	Name	Author
class	Gastropoda
subclass	Caenogastropoda	(Cox 1959)
superfamily	Epitonioidea	(Berry 1910)
family	Epitoniidae	Berry 1910
genus	Hirtoscala	Bolten 1798
species	tincta	(Carpenter 1864)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Hirtoscala tincta Carpenter 1864 [tinted wentletrap]

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Invalid names: Epitonium (Nitidiscala) eelense Durham 1937 [synonym], Epitonium (Nitidiscala) tinctum bormanni Strong 1941 [synonym], Epitonium (Nitidoscala) continuatum Oldroyd 1925 [synonym], Epitonium (Nitidoscala) subcoronatum Carpenter 1864 [synonym], Epitonium continuatum Oldroyd 1925 [synonym]

Diagnosis

Reference	Diagnosis
J. W. Durham 1937 (Epitonium (Nitidiscala) eelense)	Shell nuclear moderately convex whorls; sutures fairly deep; ornamented by 15 prominent, rather heavy varices that usually meet those of preceding whorl in the suture but do not fuse with them; some varices slightly reflexed posteriorly while others are not; a slight angle in varices just below suture makes an obscure coronation; varices ascending the spire in a line that is slightly spiral to the left; base of the body whorl slightly angulated almost forming a basal keel; aperture ovate, auricle rather prominent.

Measurements

No measurements are available

Composition:

aragonitef

Composition 2:

low Mg calcitef

Entire body:

yesg

Adult width:

10 to < 100g

Adult height:

10 to < 100g

Thickness:

thing

Folds:

noneg

Ribbing:

minorg

Spines:

noneg

Internal reinforcement:

noneg

Locomotion:

slow-movingf

Life habit:

semi-infaunalg

Diet:

carnivoref

Vision:

limitedc

Created:

2009-03-16 10:37:01

Modified:

2018-03-19 13:34:38

Source: g = genus, f = family, c = class

References: Kiessling 2004, Abbott and Dance 1986, Beu et al. 1990

Age range: base of the Messinian to the top of the Late/Upper Pleistocene or 7.24600 to 0.01170 Ma

Collections (27 total)

Time interval	Ma	Country or state	Original ID and collection number
Messinian	7.246 - 5.333	USA (California)	Epitonium eelense (194796)
Late/Upper Pliocene	3.6 - 2.58	USA (California)	Epitonium tinctum (214966 214967) Nitidiscala tincta (238622 238623)
Gelasian	2.58 - 1.8	USA (California)	Epitonium tinctum (189765)
Early/Lower Pleistocene	2.58 - 0.774	USA (California)	Epitonium tinctum (168757)
Pleistocene	2.58 - 0.0117	Mexico (Baja California)	Epitonium tinctum (96590 96592 96593)
Pleistocene	2.58 - 0.0117	USA (California)	Epitonium tinctum (201624 237610)
Middle Pleistocene	0.774 - 0.129	USA (California)	Epitonium tinctum, Epitonium continuatum, Epitonium subcoronatum (138723) Scala tincta (222663 222664)
Late/Upper Pleistocene	0.129 - 0.0117	Mexico (Baja California)	Epitonium tinctum (96591)
Late/Upper Pleistocene	0.129 - 0.0117	USA (California)	Epitonium tinctum (59063 160361 219029 219030 226458 240593 240646) Scala tincta (222659 222660 222661 222662)