PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Heyuannia yanshini

Reptilia - Oviraptoridae

Taxonomy

Ingenia yanshini was named by Barsbold (1981). Its type specimen is PST GIN AN MNP 100/30, a partial skeleton, and it is a 3D body fossil. Its type locality is Khermeen Tsav, Red Beds (PIN coll. 3142) [SMPE], which is in a Campanian/Campanian terrestrial horizon in the Baruungoyot Formation of Mongolia. It was considered monophyletic by Currie (2000).

It was recombined as Oviraptor yanshini by Paul (1988); it was recombined as Ajancingenia yanshini by Easter (2013), Wang et al. (2016), Norell et al. (2018); it was recombined as Heyuannia yanshini by Funston et al. (2018), Yun (2019), Lee et al. (2019), Funston (2020), Funston et al. (2021).

Synonymy list

Year	Name and author
1981	Ingenia yanshini Barsbold p. 35
1983	Ingenia yanshini Barsbold p. 93
1988	Oviraptor yanshini Paul p. 378
1992	Ingenia yanshini Currie p. 246
1992	Ingenia yanshini Le Loeuff et al. p. 338
1996	Ingenia yanshini Osmólska p. 33
1997	Ingenia yanshini Norell and Makovicky p. 3
1997	Ingenia yanshini Osmólska p. 41
2000	Ingenia yanshini Currie p. 438
2001	Ingenia yanshinii Norell et al. p. 13
2002	Ingenia yanshini Maryanska et al. p. 101 fig. 1
2003	Ingenia yanshini Rauhut p. 40
2004	Ingenia yanshini Lü et al. p. 109 fig. 8
2004	Ingenia yanshini Osmólska et al. p. 167
2005	Ingenia yanshini Lü and Zhang p. 417
2007	Ingenia yanshini Juárez Valieri et al. p. 61
2007	Ingenia yanshani Turner et al. p. 17
2010	Ingenia yanshini Longrich et al. p. 953 fig. 7
2010	Ingenia yanshini Xu and Han p. 12
2012	Ingenia yanshini Fanti et al. p. 1
2013	Ajancingenia yanshini Easter
2014	Ingenia yanshini Lamanna et al. p. 10 fig. 6
2015	Ingenia yanshini Funston et al. p. 186
2016	Ingenia yanshini Funston and Currie p. 10
2016	Ajancingenia yanshini Wang et al. p. 8
2018	Heyuannia yanshini Funston et al. p. 109–110
2018	Ajancingenia yanshini Norell et al. p. 13
2019	Heyuannia yanshini Lee et al. p. 2
2019	Heyuannia yanshini Yun p. 295
2020	Heyuannia yanshini Funston p. 136 fig. 20
2021	Heyuannia yanshini Funston et al. p. 7

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()

Rank	Name	Author
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Theropoda	()
—	Neotheropoda
—	Averostra	Paul 2002
—	Tetanurae
—	Coelurosauria	()
—	Maniraptora
infraorder	Oviraptorosauria
family	Oviraptoridae
subfamily	Heyuanninae
genus	Heyuannia
species	yanshini	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Heyuannia yanshini Barsbold 1981

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Diagnosis

Reference	Diagnosis
G. F. Funston et al. 2018	Small oviraptorid oviraptorosaur diagnosed by the following features: low ridge on medial side of surangular; angular contributing to retroarticular process; fused axis intercentrum; relatively large epipophyses on anterior cervical vertebrae; accessory foramen anteroventral to lateral pleurocoel of posterior cervical vertebrae; eight sacral vertebrae (shared with Ne. barsboldi); 30 caudal vertebrae with hatchet- shaped transverse processes; chevrons with distal bulb; unfused scapulocoracoid with unexpanded posterior blade, laterally-facing glenoid, and small biceps tubercle; humerus with large deltopectoral crest oc- cupying nearly half of humeral length and large ectepicondyle; ulna with large lateral distal process; metacarpal I short (two-thirds length of metacarpus) and broad; metacarpal II and III subequal in length; ungual I-2 longer than metacarpal I*; fibula contacting calcaneum; metatarsal I spade-shaped; metatarsal II longer than metatarsal IV.

Measurements

No measurements are available

Composition:

hydroxyapatiteinfrao

Entire body:

yesinfrao

Adult length:

1.0 to < 10infrao

Adult width:

0.1 to < 1.0infrao

Adult height:

1.0 to < 10infrao

Thickness:

intermediateinfrao

Architecture:

compact or denseinfrao

Ontogeny:

accretion, modification of partsinfrao

Grouping:

solitaryo

Environment:

terrestrialinfrao

Locomotion:

actively mobileinfrao

Life habit:

ground dwellinginfrao

Depth habitat:

surfaceinfrao

Diet:

omnivoreinfrao

Reproduction:

oviparousinfrao

Dispersal:

direct/internalinfrao

Dispersal 2:

mobileinfrao

Created:

2005-03-06 14:07:56

Modified:

2005-03-06 16:07:56

Source: infrao = infraorder, o = order

References: Marsh 1875, Averianov and Yarkov 2004

Age range: base of the Middle Campanian to the top of the Late/Upper Campanian or 83.50000 to 70.60000 Ma

Collections: one only

Time interval	Ma	Country or state	Original ID and collection number
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	Mongolia (Omnogov)	Oviraptor sp. (type locality: 11582)