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Gspsaurus pakistani

Reptilia - Gspsauridae

Taxonomy
Gspsaurus pakistani was named by Malkani (2014). Its type specimen is MSM-79-19, a partial skull (articulated upper and lower jaws with teeth, palatal processes, left quadrate, partial quadratojugal, possibly lowermost portion of squamosal, mandible rami and), and it is a 3D body fossil. Its type locality is Alam Kali Kakor, DL-19 (central), which is in a Maastrichtian fluvial claystone in the Vitakri Formation of Pakistan.

Synonyms
Synonymy list
YearName and author
2014Gspsaurus pakistani Malkani p. 56
2014Maojandino alami Malkani p. 169
2014Gspsaurus pakistani Malkani p. 872
2014Maojandino alami Malkani p. 872
2015Gspsaurus pakistani Malkani p. 4
2015Maojandino alami Malkani p. 6
2015Gspsaurus pakistani Malkani pp. 93, 95–96
2015Maojandino alami Malkani pp. 93, 97
2015Gspsaurus pakistani Malkani p. 251
2015Maojandino alami Malkani p. 251
2016Gspsaurus pakistani Malkani p. 129
2016Maojandino alami Malkani p. 129
2017Gspsaurus pakistani Malkani p. 117
2017Maojandino alami Malkani p. 117
2019Gspsaurus pakistani Malkani p. 628
2020Gspsaurus pakistani Malkani p. 409
2020Maojandino alami Malkani p. 409
2020Gspsaurus pakistani Malkani pp. 455, 461–463

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
Archosauromorpha(Huene 1946)
RankNameAuthor
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Saurischia()
Sauropoda()
Gravisauria
Eusauropoda
Neosauropoda
Macronaria
Titanosauriformes
Titanosauria
PoripuchiaMalkani 2020
familyGspsauridae
subfamilyGspsaurinae
genusGspsaurus
speciespakistani

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Gspsaurus pakistani Malkani 2014
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Invalid names: Maojandino alami Malkani 2014 [synonym]
Diagnosis
ReferenceDiagnosis
M. S. Malkani 2016 (Maojandino alami)a thickest, broad and long neck, short tail, heavy and stocky body
M. S. Malkani 2016The anterior dentary symphysis seems to be weak. The anterior portions of dentary rami are V-shaped. It has relatively long, narrow, slightly oval, slender and slightly recurved teeth.
M. S. Malkani 2019Gspsaurus pakistani medium sized sauropod shares with the Titanosauria as vertebrae lacking hyposphene-hypantrum articulations; procoelous caudals (except the first caudal); forward insertion of neural arches on caudals; and prominent olecranon process. Gspsaurus pakistani autapomorphies are as small sized spongy skull; teeth circular to subcircular, slender, slightly recurved and conical. The thickness of diameter decreases gradually from base to tip; palatal shelf be- tween the ventral palatal process and dorsal palatal process forms left and right maxillary canals; dorsal palatal shelf just below the left and right premaxillae and between the dorsal palatal processes forms reverse triangular premaxillary canal; some teeth cone blunted showing wear facet, broad U shaped upper teeth row and V shaped lower teeth row. Reversed V-shaped ventral palatal processes with oval shaped rod contacted week with maxilla; V-shaped small dorsal palatal processes attached on the contact of maxilla and premaxilla. Dentary ramus anterior depth is slightly less than mid length, dentary with long anteroposterior symphyses, angled 15° or more anteriorly to axis of jaw ramus, transversely thick and anteroposteriorly lense shaped proximal tibia and transversely oval shaped distal tibia.
M. S. Malkani 2020Gspsaurus pakistani a medium sized and stocky sauropod dinosaur sharing with the Titanosauria as vertebrae lacking hyposphene-hypantrum articulations, procoelous caudals (except first biconvex caudal), forward insertion of neural arches on caudals, and prominent olecranon process on ulna. Gspsaurus pakistani a medium sized and stocky sauropod dinosaur sharing with the Poripuchia as the distal and also distalmost caudals are procoelous besides the anterior and middle procoelous caudals which is the character of Lithostrotian.

Gspsaurus pakistani autapomorphies are as small sized spongy skull (but larger than Saraikimasoom). Teeth are circular to subcircular, slender, slightly recurved and conical like-the thickness of diameter decreases gradually from base to tip (while the Pakisaurus and Rapetosaurus have teeth with almost constant thickness from base to tip but except tip). Teeth are longer than Saraikimasoom teeth. Teeth slender indices vary from 3–5 (while Pakisaurus and Rapetosaurus teeth have Slender Indices more than 5). Some teeth are cone blunted showing wear facet, some teeth tips are rounded and a few have pointed tip. Palatal shelf process is deep and has about 45° limb inclination (while Saraikimasoom have shallow inclination upto 25°). Palatal shelf between the ventral palatal process and dorsal hook process form left and right maxillary canals. Dorsal palatal processes formed hook with its limbs contacted on the suture of maxilla and premaxilla. Dorsal hook limb processes form reverse triangular premaxillary canal. Broad U shaped upper teeth row and V shaped lower teeth row (while Saraikimasoom have both lower and upper jaws as U-shaped teeth rows). Reversed gentle V-shaped ventral palatal processes with expanded rectangle shaped distal ends which are contacted with the maxilla. V-shaped small dorsal palatal processes or hook limb attached on the suture of maxilla and premaxilla. Dentary have long anteroposteriorly symphysis (while dentary with narrow anteroposteriorly symphysis found in Saraikimasoom). Anterior dentary is rounded and has no chin. Dentary symphysis is an extension of dentary ramus in the almost same direction (while it is perpendicular to dentary ramus axis in Saraikimasoom). Dentary ramus anterior depth is slightly less than mid length (while Saraikimasoom have anterior dentary with slightly more depth than mid dentary). Dentary with long anteroposteriorly symphysis is angled 15° or more anteriorly to axis of jaw ramus (while Saraikimasoom have high angle anterior profile of dentary symphysis). Large braincase with subrectangle shaped occipital condyle with sagital dorsoventral prominent groove (while Saraikimasoom have relatively small braincase with feeble sagital groove, Pakisaurus and probably Isisaurus have D-shaped occipital condyle). First caudal is biconvex and broad with relatively less long than Saraikimasoom. Caudals especially mid caudals are slightly ventrally reduced upto ratio 1.5 (while Pakisaurus and Isisaurus have not ventrally reduced caudals, however in Saraikimasoom ventral reduction is prominent and ratio vary from 1.5 to 2). Trirays robust procoelous distalmost caudal (while Sa- raikimasoom have relatively less robust trirays procoelous distalmost caudal, Pakisaurus have horizontal groove in the mid of distalmost caudal). Distal scapula is quite expanded transversely than the distal scapula of Saraikimasoom, Pakisaurus and Isisaurus. Acetabulum has anteroposteriorly elongated fibrous structures. Acetabulum has D-shaped thick and long condyle or peduncle for pubis and relatively small and thin condyle or peduncle for pubis. Rectangle or subrectangle shaped cross section of shaft just below the femur head with transversely long and anteroposteriorly relatively less broad than Saraikimasoom (while the Saraikimasoom have more robust rectangle shaped cross section with thicker anteroposteriorly). Transversely thick biconvex lense shaped proximal tibia which anteroposterior width is slightly more than transverse width (while Saraikimasoom have subsquare shaped proximal tibia with equal transverse and anteroposterior widths, Pakisaurus and Isisaurus have flattened and transversely compressed proximal tibia). Distal tibia is transversely quite thick and quite expanded than Pakisaurus and Isisaurus (while Pakisaurus and Isisaurus have anteroposteriorly broad distal tibiae). The expanding nature of tibia shows close articulation with fibula (while relatively far articulation found in fibula and tibia when articulated). Proximal fibula orientation shifted the trend of proximal fibula from anteroposterior to almost mediolateral due to transversely expanded fibular condyle of proximal tibia (instead of anteroposterior or anteromedial to posterolateral orientation as in Pakisaurus and Isisaurus) when articulated with proximal tibia. Two types of armour bones and osteoderms are found, the first small rectangular mosaic type is relatively thin plate (Saraikimasoom have more thick armour mosaic plate) and secondly the large oval ellipsoidal plate with median cut or groove like Malawisaurus (while Saraikimasoom have ellipsoidal subcircular plate without median groove or cut).
Measurements
No measurements are available
Composition: hydroxyapatiteinfrao
Entire body: yesinfrao
Adult length: 10 to < 100infrao
Adult width: 10 to < 100infrao
Adult height: 10 to < 100infrao
Thickness: thickinfrao
Architecture: compact or denseinfrao
Form: sphericalinfrao
Ontogeny: accretion, modification of partsinfrao
Grouping: gregariousinfrao
Environment: terrestrialinfrao
Locomotion: actively mobileinfrao
Life habit: ground dwellinginfrao
Diet: herbivoreinfrao
Reproduction: oviparousinfrao
Dispersal: direct/internalinfrao
Dispersal 2: mobileinfrao
Created: 2004-12-14 12:49:10
Modified: 2004-12-14 14:49:10
Source: infrao = infraorder
Reference: Marsh 1875

Age range: Late/Upper Maastrichtian or 70.60000 to 66.00000 Ma

Collections (6 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Maastrichtian70.6 - 66.0Pakistan (Balochistan) Gspsaurus pakistani (79428 79429 79664 79665 79668) Maojandino alami, Marisaurus jeffi (type locality: 79666)