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Helicina

Gastropoda - Heterostropha

Synonymy list
YearName and author
2017Helicina Bouchet et al. p. 361

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Spiralia
superphylumLophotrochozoa
phylumMollusca
classGastropoda
orderHeterostropha()
RankNameAuthor
infraclassEuthyneura(Spengel 1881)
—TectipleuraSchrödl et al. 2011
—PanpulmonataJörger et al. 2010
superorderEupulmonata(Haszprunar and Huber 1990)
suborderGeophila(Férussac 1819)
orderStylommatophora(Schmidt 1856)
suborderHelicina
suborderHelicina

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subor. Helicina
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Infraor. Clausilioidei
+
Superfm. Clausilioidea Gray 1855
+
Fm. Clausiliidae Gray 1855
+
Subfm. †Alopiinae Wagner 1913
+
G. †Cochlodina Férussac 1821
Subfm. †Clausiliinae Gray 1855
+
G. †Clausilia Draparnaud 1805
G. †Truciella Nordsieck 1978
G. †Constricta Boettger 1877
G. Macrogastra Hartmann 1841
G. †Pseudidyla Boettger 1877
+
Subg. †Pseudidyla (Canaliciella) Nordsieck 1981
†Pseudidyla martingrossi Harzhauser and Binder 2008
†Pseudidyla moersingensis Boettger 1877
†Pseudidyla schultzi Binder 2004
G. †Regiclausilia Nordsieck 1981
+
†Regiclausilia patula Nordsieck 1981
G. †Serrulastra Nordsieck 1981
+
Subg. †Serrulastra (Serrulastra) Nordsieck 1981
Subg. †Serrulastra (Serruplica) Nordsieck 1981
Subfm. †Serrulininae Ehrmann 1927
+
G. †Nordsieckia Truc 1972
G. †Serrulella Nordsieck 1978
G. Triloba Vest 1867
+
†Triloba pappi Schnabel 2012
G. †Trolliella Nordsieck 1981
+
†Trolliella silesiaca Nordsieck 1981
Fm. †Filholiidae Wenz 1923
+
G. †Triptychia Sandberger 1876
+
†Triptychia ederae Binder 2004
†Triptychia kleini Schnabel 2006
+
Invalid names: Clausilia grandis Klein 1846 [synonym]
†Triptychia margaretae Andreae 1904
†Triptychia randeckiana Kranz 1908
†Triptychia suevica Sandberger 1875
Invalid names: Palaeostoidae Nordsieck 1986 [empty]
Infraor. Helicoidei
+
Superfm. Helicoidea Rafinesque 1815
+
Fm. Camaenidae Pilsbry 1895
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G. †Amphidromus Albers 1850
G. Labyrinthus Beck 1837 [land snail]
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†Labyrinthus obtusus Anderson and Hanna 1925
G. †Pleurodonte Fischer and Waldheim 1807
+
Subg. Pleurodonte (Galactochilus) Sandberger 1875
Subg. Pleurodonte (Pleurodonte) Fischer von Waldheim 1807
†Pleurodonte michalkovaci Binder and Harzhauser 2008
†Pleurodonte norica Binder 2004
G. Zachrysia Pilsbry 1894 [land snail]
+
†Zachrysia fraterna Roth 1988
Fm. Elonidae Gittenberger 1979
+
Subfm. Klikiinae Nordsieck 1986
+
G. Apula Boettger 1909
G. Klikia Pilsbry 1895
G. Pseudochloritis Boettger 1909
+
+
Invalid names: Helix inflexa Klein 1847 [synonym], Oxychilus subnitens Seemann 1926 [synonym]
Fm. Geomitridae Boettger 1909
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G. Cernuella Schlüter 1838
G. Trochoidea Brown 1827
+
G. Xerosecta Monterosato 1892
Fm. Helicidae Rafinesque 1815
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Subfm. Ariantinae Mörch 1864
+
G. †Agalactochilus Kadolsky et al. 2016
G. Helicigona Risso 1826
G. Cepaea Held 1838
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†Cepaea etelkae Halavats 1923
†Cepaea eversa Deshayes 1851
†Cepaea sylvestrina Schlotheim 1820
G. Eremina Pfeiffer 1855
G. Helix Linnaeus 1758
+
Subg. Helix (Iberus) Montfort 1910
Subg. Helix (Macularia) Albers 1850
Subg. Helix (Patula) Held 1838
Subg. †Helix (Tachea) Turton 1831
Subg. †Helix (Tacheocampylaea) Pfeiffer 1877
Subg. Helix (Trigonostoma) Fitzinger 1833
†Helix desmarestina Brongniart 1810
†Helix dubia Deshayes 1824
†Helix evanstonensis White 1878
†Helix ferrantii Deshayes 1824
†Helix lemani Brongniart 1810
†Helix moroguesi Brongniart 1810
†Helix tristani Brongniart 1810
G. Marmorana Hartman 1844
+
†Marmorana serpentina Ferussac 1826
G. Massylaea Möllendorff 1898
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Massylaea massylaea Morelet 1851
Massylaea vermiculata Müller 1774
G. Megalotachea Pfeffer 1930
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†Megalotachea turonensis Deshayes 1831
G. Otala Schumacher 1817
G. Palaeotachea Jooss 1912
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†Palaeotachea renevieri Maillard 1892
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Invalid names: Cepaea dentula Ehrat and Jooss 1921 [synonym], Cepaea silvana Wenz 1923 [synonym]
G. Tacheocampylaea Pfeiffer 1877
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†Tacheocampylaea raspaili Payraudeau 1826
Fm. Helicodontidae Kobelt 1904
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G. Helicodonta Ferussac 1821
G. Protodrepanostoma Germain 1929
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Fm. Hygromiidae Tryon 1866
+
G. Caracollina Beck 1837
G. †Helicopsis Fitzinger 1833
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†Helicopsis piedmontanica Harzhauser et al. 2015
G. Leucochroopsis Boettger 1908
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Subfm. Monachainae Wenz 1930
+
G. †Euomphalia Westerlund 1889
G. †Pseudoxerotricha Boettger 1911
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G. Trochulus Chemnitz 1786
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Invalid names: Trichia Hartmann 1840 [synonym]
G. Urticicola Lindholm 1927
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†Urticicola perchtae Salvador 2013
Fm. Pleurodontidae Ihering 1912
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G. Ganesella Blanford 1863
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Ganesella martini Oostingh 1935
Fm. Polygyridae Pilsbry 1895
+
G. Daedalochila Beck 1837
G. †Euchemotrema Archer 1939
+
†Euchemotrema occidaneum Roth and Emberton 1994
G. Patera Albers 1850
G. Polygyra Say 1818
G. Triodopsis Rafinesque 1819
Fm. Sphincterochilidae Zilch 1960
+
G. Sphincterochila Ancey 1887
Fm. Trissexodontidae Nordsieck 1987
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G. Praeoestophorella Pfeffer 1930
+
+
Invalid names: Caracollina phacodes Rasser 2013 [synonym], Oestophora phacodes Ehrat and Jooss 1921 [synonym]
Superfm. Sagdoidea Pilsbry 1895
+
Fm. Sagdidae Pilsbry 1895
+
G. Microphysula Cockerell 1926
Infraor. Limacoidei
+
Superfm. Gastrodontoidea Tryon 1866
+
Fm. Gastrodontidae Tryon 1866
+
G. †Janulus Lowe 1852
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†Janulus austriacus Harzhauser and Binder 2004
†Janulus spadinii Manganelli et al. 2011
†Janulus supracostatus Sandberger 1873
Fm. Oxychilidae Hesse 1927
+
G. Aegopinella Lindholm 1926
+
†Aegopinella denudata Reuss 1849
†Aegopinella depressula Harzhauser and Neubauer 2018
†Aegopinella reussi Hoernes 1856
G. Daudebardia Hartmann 1821
+
†Daudebardia praecursor Andreae 1902
Subfm. Oxychilinae Hesse 1927
+
Invalid names: Helicellinae Adams and Adams 1855 [replaced]
G. †Oxychilus Fitzinger 1833
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†Oxychilus procellarium Jooss 1918
G. Perpolita Baker 1928
+
†Perpolita miocaenica Andreae 1902
Fm. Pristilomatidae Cockerell 1891
+
G. †Eurocystina Harzhauser and Neubauer 2018
+
†Eurocystina nordsiecki Harzhauser and Neubauer 2018
G. Vitrea Fitzinger 1833
+
†Vitrea angustaeumbilicata Harzhauser and Neubauer 2018
†Vitrea procrystallina Andreae 1902
Superfm. Helicarionoidea Bourguignat 1877
+
G. Trochozonites Pfeffer 1883
+
Trochozonites arabica Neubert and van Damme 2012
Superfm. Limacoidea Lamarck 1801
+
Fm. †Agriolimacidae Wagner 1935
+
G. Deroceras Rafinesque 1820
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†Deroceras aenigma Leonard 1950
Fm. Limacidae Lamarck 1801
+
G. Lehmannia Heynemann 1863
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†Lehmannia excavata Andreae 1904
G. Limax Linnaeus 1758
+
†Limax fossilis Sacco 1896
G. †Mennoia Harzhauser and Neubauer 2018
+
†Mennoia sculpturata Harzhauser and Neubauer 2018
G. †Neubertella Harzhauser and Neubauer 2018
+
†Neubertella pulchra Harzhauser and Neubauer 2018
Fm. Vitrinidae Fitzinger 1833
+
G. †Oligolimax Fischer 1878
G. Phenacolimax Stabile 1859
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†Phenacolimax crassitesta Andreae 1902
G. Semilimax Agassiz 1845
G. Vitrina Draparnaud 1801
+
†Vitrina obesa Pan 1977
†Vitrina obliqua Pan 1977
Vitrina suevica Sandberger 1876
Superfm. Parmacelloidea Fischer 1856
+
Fm. Milacidae Ellis 1926
+
G. Milax Gray 1855
+
†Milax oppoliensis Andreae 1904
Fm. Parmacellidae Fischer 1856
+
G. †Parmacella Cuvier 1804
Superfm. Zonitoidea Morch 1864
+
Fm. Zonitidae Mörch 1864 [true glass snail]
+
G. Aegopis Fitzinger 1833
Subfm. †Archaeozonitinae Pfeiffer 1930
+
G. †Archaeozonites Sandberger 1873
G. Glyphyalinia Martens 1892
G. Hawaiia Gude 1911
G. Miozonites Pfeffer 1929
+
Miozonites costatus Binder 2002
G. Nesovitrea Cooke 1921
+
†Nesovitrea boettgeriana Clessin 1877
†Nesovitrea disciformis Lueger 1981
G. Zonitoides Lehmann 1862
+
Subg. Zonitoides (Zonitoides) Lehmann 1862
†Zonitoides schaireri Schlickum 1978
Infraor. Oleacinoidei
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Superfm. Oleacinoidea Adams and Adams 1855
+
Fm. Oleacinidae Adams and Adams 1850
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G. Oleacina Röding 1798
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Subg. Oleacina (Salasiella) Strebel 1878
G. †Pseudoleacina Wenz 1914
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†Pseudoleacina eburnea Klein 1853
†Pseudoleacina fossilis Andreae 1902
G. †Spiraxis Adams 1850
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†Spiraxis tampae Dall 1915
Fm. Spiraxidae Baker 1939
+
G. Euglandina Crosse and Fischer 1870
Subfm. Euglandininae Baker 1941
+
G. †Palaeoglandina Wenz 1914
Infraor. Orthalicoidei
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Superfm. Orthalicoidea Albers 1860
+
Fm. Bulimulidae Tryon 1867
+
Subfm. Bulimulinae Tryon 1867
+
G. Bulimulus Leach 1814
G. †Oreoconus McKenna et al. 1962
+
†Oreoconus planispira McKenna et al. 1962
G. †Tocobaga Auffenberg et al. 2015
+
Tocobaga americanus Heilprin 1886
†Tocobaga floridanus Conrad 1846
†Tocobaga wakullae Mansfield 1937
Fm. Odontostomidae Pilsbry and Vanatta 1898
+
G. Bahiensis Jousseaume 1877
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†Bahiensis priscus Cabrera and Martínez 2012
G. Plagiodontes Doering 1876
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†Plagiodontes dentatus Wood 1828
Fm. Orthalicidae Albers 1860
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Subfm. Orthalicinae Albers 1860
+
G. Orthalicus Beck 1837
Invalid names: Vidaliellidae Nordsieck 1986 [empty]
Superfm. Plectopyloidea Möllendorff 1898
+
Fm. Plectopylidae Möllendorff 1898
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G. †Plectopylis Benson 1860
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†Plectopylis degerbolae Solem 1966
Superfm. Punctoidea Morse 1864
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Fm. Charopidae Hutton 1884
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Subfm. Charopinae Hutton 1884
+
G. Lagivala Solem 1982
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†Lagivala davidi Ladd 1968
G. Vatusila Solem 1982
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G. Colhueconus Miquel and Bellosi 2010
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†Colhueconus simpsoni Miquel and Bellosi 2010
G. Gyrocochlea Hedley 1924
G. Lilloiconcha Weyrauch 1965
G. Radiodiscus Pilsbry 1906
G. Rotadiscus Pilsbry 1926
G. Stephadiscus Hylton Scott 1981
G. Stephanoda von Martens 1860
+
†Stephanoda mazzonii Miquel and Bellosi 2004
G. Zilchogyra Weyrauch 1965
Fm. Endodontidae Pilsbry 1895
+
G. Cookeconcha Solem 1976
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†Cookeconcha antiquus Solem 1977
G. Minidonta Solem 1976
+
†Minidonta inexpectans Ladd 1958
G. †Protoendodonta Solem 1977
+
†Protoendodonta laddi Solem 1977
G. Ptychodon Ancey 1888
Fm. †Helicodiscidae Pilsbry 1927
+
G. Helicodiscus Morse 1864
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†Helicodiscus roemeri Andreae 1902
G. †Lucilla Lowe 1852
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†Lucilla miocaenica Harzhauser et al. 2015
†Lucilla subteres Clessin 1877
Fm. Oreohelicidae Pilsbry 1939
+
G. Oreohelix Pilsbry 1904 [mountain snail]
+
Oreohelix megarche Cockerell and Henderson 1912
G. Radiocentrum Pilsbry 1905
Fm. Punctidae Morse 1864
+
G. †Punctum Morse 1864
+
†Punctum parvulum Gottschick 1920
†Punctum propygmaeum Andreae 1904
Infraor. Pupilloidei
+
Superfm. Pupilloidea Turton 1831
+
Fm. Argnidae Hudec 1965
+
G. †Argna Cossmann 1889
+
†Argna oppoliensis Andreae 1902
†Argna proexcessiva Sacco 1889
Fm. Azecidae Watson 1920
+
G. Azeca Fleming 1828
+
†Azeca frechi Andreae 1902
†Azeca lubricella Boettger 1870
†Azeca pumila Slavik 1869
Fm. Chondrinidae Steenberg 1925
+
G. Granaria Held 1837
+
†Granaria grossecostata Gottschick and Wenz 1919
†Granaria moedlingensis Harzhauser and Binder 2004
†Granaria schlosseri Cossmann 1908
+
Invalid names: Pupa (Vertigo) oviformis Schlosser 1907 [replaced]
Fm. Cochlicopidae Pillsbry 1900
+
Subfm. Cochlicopinae Pilsbry 1900
+
G. Cochlicopa Ferussac 1821
G. Hypnophila Bourguignat 1858
+
†Hypnophila loxostoma Klein 1853
+
Invalid names: Cochlicopa subrimata Rasser 2013 [synonym]
Fm. Enidae Woodward 1903
+
G. Ena Turton 1831
G. †Palaeomastus Nordsieck 2014
Invalid names: Buliminusidae Kobelt 1880 [replaced]
Fm. Gastrocoptidae Pilsbry 1918
+
G. †Balticopta Balashov and Perkovsky 2020
+
†Balticopta gusakovi Balashov and Perkovsky 2020
G. Gastrocopta Wollaston 1878
+
Subg. Gastrocopta (Albinula) Sterki 1892
Subg. Gastrocopta (Sinalbinula) Pilsbry 1916
†Gastrocopta acuminata Klein 1846
†Gastrocopta nouletiana Dupuy 1850
Fm. Lauriidae Steenberg 1925
+
G. Leiostyla Lowe 1852
+
Subg. Leiostyla (Leiostyla) Lowe 1852
†Leiostyla piserai Harzhauser and Neubauer 2018
Fm. Orculidae Pilsbry 1918
+
G. †Nordsieckula Hari and Harzhauser 2014
+
†Nordsieckula falkneri Hausdorf 1995
G. †Pagodulina Clessin 1876
Fm. Partulidae Pilsbry 1900
+
G. Partula Férussac 1821
Fm. Pleurodiscidae Wenz 1923
+
G. †Pleurodiscoides Nordsieck 2014
+
Subg. †Pleurodiscoides (Pleurodiscoides) Nordsieck 2014
Fm. Pupillidae Turton 1831
+
G. †Albertanella Russell 1931
+
†Albertanella minuta Russell 1931
G. Gibbulinopsis Germain 1919
+
G. †Microstele Boettger 1886
G. Pupoides Pfeiffer 1854
+
Subg. Pupoides (Ischnopupoides) Pilsbry 1926
Fm. Pyramidulidae Kennard and Woodward 1914
+
G. Pyramidula Fitzinger 1833
+
Pyramidula cronkhitei Newcomb 1865
†Pyramidula lecontei Stearns 1902
†Pyramidula ralstonensis Cockerell 1914
Fm. Strobilopsidae Wenz 1915
+
G. †Eostrobilops Pilsbry 1908
+
†Eostrobilops boettgeri Andreae 1902
G. Strobilops Pilsbry 1893
+
Subg. †Strobilops (Strobilops) Pilsbry 1893
†Strobilops costatus Clessin 1877
+
Invalid names: Strobilops bilamellatus Clessin 1885 [synonym]
†Strobilops romani Wenz 1915
†Strobilops tiarula Sandberger 1886
Subfm. Truncatellinidae Steenberg 1925
+
G. †Truncatellina Lowe 1852
+
†Truncatellina lentilii Miller 1900
†Truncatellina pantherae Harzhauser and Neubauer 2014
†Truncatellina suprapontica Wenz and Edlauer 1942
Fm. Valloniidae Morse 1864
+
G. Vallonia Risso 1826
+
†Vallonia lepida Reuss 1849
†Vallonia subcyclophorella Gottschick 1911
†Vallonia subpulchella Sandberger 1874
Fm. Vertiginidae Fitzinger 1833
+
G. †Negulus Boettger 1889
+
†Negulus gracilis Gottschick and Wenz 1919
†Negulus suturalis Sandberger 1858
Subfm. †Nesopupinae Steenberg 1925
+
G. †Negulopsis Nordsieck 2014
Subfm. Vertigininae Fitzinger 1833
+
G. Vertigo Muller 1774
Invalid names: Achatinellidae Gulick 1873 [empty], Achatinelloidea Gulick 1873 [synonym], Cerastidae Wenz 1923 [empty], Cochlicopoidea Pilsbry 1900 [synonym], Draparnaudiidae Solem 1962 [empty], Enoidea Woodward 1903 [synonym], Partuloidea Pilsbry 1900 [synonym]
Invalid names: Orthurethra Pilsbry 1900 [synonym]
Infraor. Rhytidoidei
+
Superfm. Rhytidoidea Pilsbry 1893
+
Fm. Acavidae Pilsbry 1895
+
G. †Pebasiconcha Wesselingh and Gittenberger 1999
+
†Pebasiconcha immanis Wesselingh and Gittenberger 1999
Fm. Dorcasiidae Connolly 1915
+
G. Dorcasia Gray 1838
Fm. Megomphicidae Baker 1930
+
G. Glyptostoma Bland and Binney 1858
+
Fm. Strophocheilidae Pilsbry 1902
+
G. Austroborus Parodiz 1949
G. †Eoborus Klappenbach and Olazarri 1970
+
†Eoborus rotundus Salvador and Simone 2012
Invalid names: Acavoidea Pilsbry 1895 [synonym]
Infraor. Succineoidei
+
Superfm. Athoracophoroidea Fischer 1883
+
Fm. Athoracophoridae Fischer 1883
+
Invalid names: Athoracophorinae Fischer 1883 [empty]
Superfm. Succineoidea Beck 1837
+
Fm. Succineidae Beck 1837 [amber snail]
+
G. Oxyloma Westerlund 1885
+
†Oxyloma minima Klein 1853
Subfm. Succineinae Beck 1837
+
G. Succinea Draparnaud 1801
Invalid names: Catinellinae Odhner 1950 [empty]
Invalid names: Elasmognatha Morch 1864 [synonym]
Superfm. Testacelloidea Gray 1840
+
Fm. Testacellidae Gray 1840
+
G. †Testacella Draparnaud 1801
+
†Testacella schuetti Schlickum 1967
†Testacella zellii Klein 1853
Superfm. Urocoptoidea Pilsbry 1898
+
Fm. Cerionidae Pilsbry 1901
+
G. †Brasilennea Maury 1935
+
†Brasilennea guttula Salvador and Simone 2012
Fm. Urocoptidae Pilsbry 1898
+
G. Holospira Martens 1860
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: aragonitec
Locomotion: actively mobilec
Life habit: epifaunalc
Vision: limitedc
Created: 2004-02-29 11:40:59
Modified: 2010-02-11 01:35:02
Source: c = class
Reference: Kiessling 2004

Age range

Maximum range based only on fossils: base of the Barremian to the top of the Holocene or 129.40000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 125.0 Ma

Collections (331 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Barremian129.4 - 125.0Japan (Ishikawa) Pupilloidea sp. (32324)
Early/Lower Albian - Early/Lower Cenomanian112.03 - 93.5Argentina (Santa Cruz) Patera sp. (149928)
Late/Upper Campanian - Early/Lower Maastrichtian83.5 - 66.0USA (California) Bulimulus sp. (44268)
Maastrichtian72.1 - 66.0Uruguay Bulimulus klappenbachi (231033) Clausiliidae indet., Pupoides gnocco, Bahiensis priscus, Eoborus charruanus (231037) Succineidae indet., Bulimulus klappenbachi, Bulimulus frenguellii, Pupoides gnocco, Bahiensis priscus, Eoborus charruanus (231032) Succineidae indet., Bulimulus klappenbachi, Pupoides gnocco, Bahiensis priscus (231034) Succineidae indet., Bulimulus klappenbachi, Pupoides gnocco, Eoborus charruanus (231035) Succineidae indet., Pupoides gnocco (231036)
Maastrichtian72.1 - 66.0USA (Texas) Patera romeri (205510)
Early/Lower Maastrichtian70.6 - 66.0Romania (Hunedoara) Pyramidula grigorescui (67818)
Early/Lower Maastrichtian70.6 - 66.0Argentina (Santa Cruz) Bulimulidae indet., Holospira sp. (206969)
Early/Lower Maastrichtian70.6 - 66.0Mexico (Guerrero) Holospira sp. (7461)
Danian66.0 - 61.6USA (Texas) Holospira fallsensis (205466)
Paleocene66.0 - 56.0Austria (Salzburg) Clausilia inopinata (84335)
Paleocene66.0 - 56.0China (Yunnan) Cochlicopa headonensis, Succinea peregrina, Vitrina obliqua, Vitrina obesa (165460)
Ypresian56.0 - 47.8Brazil Plagiodontes dentatus, Brasilennea guttula, Eoborus rotundus (231038)
Eocene56.0 - 33.9USA (California) Helminthoglypta obtusa (183453)
Eocene56.0 - 33.9USA (Wyoming) Vertigo arenula, Vertigo atavuncula (197777)
Wasatchian55.8 - 50.3USA (Wyoming) Discus ralstonensis (15779) Holospira sp. (15759 15780 15832 27353) Oreohelix sp., Pupillidae indet., Polygyra sp. (15793)
Early/Lower Eocene55.8 - 48.6USA (Wyoming) Hendersonia evanstonensis, Oreohelix megarche, Microphysula sp., Discus ralstonensis, Albertanella minuta, Grangerella sinclairi (15081)
Bridgerian50.3 - 46.2USA (Wyoming) Gastrocopta sp., Oreoconus sp. (16296) Helix wyomingensis (106325) Holospira sp. (16431) Oreoconus planispira (16423) Oreoconus sp. (191230) Oreoconus sp., Holospira sp. (16244) Vertigo arenula (191226 191227)
Mustersan48.0 - 42.0Argentina (Chubut) Rotadiscus sp., Helix dentata, Colhueconus simpsoni, Gyrocochlea sp., Lilloiconcha sp., Radiodiscus sp., Stephanoda mazzonii, Stephadiscus sp., Zilchogyra sp. A, Zilchogyra sp. B (105837)
Lutetian47.8 - 41.3Namibia (Sperrgebiet) Trigonephrus sp., Dorcasia sp. (149173)
Lutetian47.8 - 41.3USA (California) Zachrysia fraterna (16712)