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Taxonomy
Neocalmonia (Quadratispina) was named by Gandl (1972). Its type is Neocalmonia (Quadratispina) quadratispinosa.
It was reranked as Quadratispina by Van Viersen and Vanherle (2018).
It was assigned to Neocalmonia by Gandl (1972); and to Asteropyginae by Van Viersen and Vanherle (2018).
It was reranked as Quadratispina by Van Viersen and Vanherle (2018).
It was assigned to Neocalmonia by Gandl (1972); and to Asteropyginae by Van Viersen and Vanherle (2018).
Subtaxa
Synonymy list
| Year | Name and author |
|---|---|
| 1972 | Neocalmonia (Quadratispina) Gandl |
| 2018 | Quadratispina Van Viersen and Vanherle |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Quadratispina Gandl 1972
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†Neocalmonia (Quadratispina) quadratispinosa Gandl 1972
†Quadratispina asturica Haas 1970
†Quadratispina excelsa Van Viersen and Vanherle 2018
†Quadratispina hispanica Richter and Richter 1926
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| A. P. Van Viersen and W. Vanherle 2018 | Occipital ring bearing a median tubercle. Pygidial axis comprising 10-13 (+1) narrow (sag.), sharply demarcated rings most of which bearing a median tubercle (weak or absent in posterior rings). 5-6 pairs of pleurae with a strongly modified supradevonicus pattern: these consist of steeply inclined posterior bands and moderately narrow (exsag.) yet sharply demarcated and individually vaulted anterior bands. The abaxial half of the posterior pleural band is often widened; in those cases it overhangs the anterior band of the posteriorly lying pleura; further abaxially the posterior pleural bands are extended as roof-shaped swellings on the pygidial border and into pleural lappets; border furrows at these transitional points nonetheless visible (indented). Interpleural furrows narrow and sharp. Large fenestrae may be developed. Pleural furrows equivalent to average width (exsag.) of a posterior pleural band; considerably deepened. Pleural lappets shorter than first pleural pair, lumpish, sharply keeled, and as a result subquadratical in section, always posteriorly directed, and at least the fifth pair convergent. Terminal lappet about as long as fifth pleural lappet; triangular. |
Measurements
No measurements are available
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| Source: f = family, subo = suborder, c = class | |||||
| References: Whittington et al. 1997, Fortey and Owens 1999, Aberhan et al. 2004 | |||||
Age range: base of the Early/Lower Frasnian to the top of the Givetian or 383.70000 to 382.70000 Ma
Collections (4 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Givetian | Spain (Aragon) | Neocalmonia quadratispinosa (40446) | |
| Early/Lower Frasnian | Belgium | Q. excelsa (234782) | |
| Early/Lower Frasnian | Spain (Castille & Léon) | Heliopyge asturica (234789 234791) |