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Paraortygoides argillae
Taxonomy
Paraortygoides argillae was named by Mayr and Kitchener (2024). Its type specimen is NMS.Z.2021.40.174, a set of limb elements (roximal end of left femur, distal ends of both femora, distal portion of right tibiotarsus, proximal and distal portions of both tarsometatarsi, pedal phalanges), and it is a 3D body fossil. Its type locality is Walton-on-the-Naze Division A2 (NMS), which is in a Ypresian marine horizon in the London Clay Formation of the United Kingdom.
Synonymy list
| Year | Name and author |
|---|---|
| 2024 | Paraortygoides argillae Mayr and Kitchener p. 2 figs. Figs. 1, 2 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Paraortygoides argillae Mayr and Kitchener 2024
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Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| G. Mayr and A. C. Kitchener 2024 | Smaller than Argillipes aurorum, “A.” paralectoris, Percolinus venablesi, and “Paraortygoides” radagasti. Distinguished from Paraortygoides messelensis Mayr, 2000 (Gallinuloididae) in that first phalanx of second toe proportionally shorter (slightly shorter than the first phalanx of third toe in P. messelensis, but somewhat longer in ?Paraortygoides argillae, sp. nov.; Fig. 2LL, NN) and in that ungual phalanges straighter and proportionally longer. Apart from a smaller size the new species differs from “Paraortygoides” radagasti Dyke and Gulas, 2002 in that trochlea metatarsi II proportionally shorter and hypotarsus with closed canal. Distinguished from Gallinuloides
wyomingensis Eastman, 1900 (Gallinuloididae) in that tarsometatarsus with proportionally shorter dorsal portion of articular surface of trochlea metatarsi III (proximal terminus at level of distal end of trochlea metatarsi II, whereas it reaches to middle of trochlea metatarsi II in Gallinuloides; Fig. 2AA, BB), in that first phalanx of second toe proportionally shorter (slightly shorter than first phalanx of third toe in G. wyomingensis but slightly longer in ?P. argillae; Fig. 2LL, MM), and in that distal end of trochlea metatarsi IV more laterally slanted. Differs from the somewhat larger Argillipes aurorum Harrison and Walker, 1977 (fam. inc. sed.) in that hypotarsus with proportionally smaller canal for tendon of musculus flexor digitorum longus (Fig. 2FF, GG); the distal end of the tarsometatarsus differs from a distal tarsometatarsus that was considered by Mayr and Smith (2019) to possibly be from A. aurorum in that the trochleae are more splayed. Distinguished from Percolinus venablesi Harrison and Walker, 1977 (fam. inc. sed.) in that tuberositas musculi tibialis cranialis less pronounced and in that hypotarsus proximodistally shorter (Fig. 2D, J). Differs from “Argillipes” paralectoris and “Percolinus” proudlocki in that hypotarsus with canal for tendon of musculus flexor digitorum longus (this canal is absent in “A.” paralectoris and “P.” proudlocki; Fig. 2JJ). Differentiated from Coturnipes cooperi Harrison and Walker, 1977 (fam. inc. sed.) in that trochlea metatarsi IV more laterally splayed, incisura intertrochlearis lateralis wider, and plantar articular surface of trochlea metatarsi III less asymmetric (Fig. 2Q–W). Distinguished from Litoripes medius Harrison and Walker, 1979 (fam. inc. sed.) in that tarsometatarsus with proportionally wider distal end and trochlea metatarsi IV reaching farther distally. The proximal tarsometatarsus referred to B. transitoria by Zelenkov (2021) differs from that of the new species in being smaller and in that crista medialis hypotarsi not as strongly plantarly protruding. ?Paraortygoides argillae, sp. nov. is distinguished from Bumbanipodius magnus Zelenkov, 2021 (fam. inc. sed.) in that tarsometatarsus without laterally prominent tubercle for ligamentum collaterale mediale. It can be differentiated from the species of Quercymegapodius Mourer-Chauviré, 1992 (Quercymegapodiidae) in that foramen vasculare distale situated farther proximally, plantar articular surface of trochlea metatarsi proximodistally longer than mediolaterally wide (about the same length and width in Quercymegapodius), and in that trochleae metatarsorum II and IV proportionally longer. Differs from Ameripodius alexis Mourer-Chauviré, 2000 (Quercymegapodiidae) in that hypotarsus with intermediate crest less developed (see Zelenkov, 2021: fig. 2l), and in that foramen vasculare distale situated farther proximally (comparisons with Ameripodus silvasantosi Alvarenga, 1995 are not possible owing to a lack of overlap in the known skeletal elements). Distinguished from the species of the Paraortygidae in that trochleae metatarsorum of tarsometatarsus more splayed. Differs from Chambiortyx cristata Mourer-Chauviré et al., 2013, in that trochlea metatarsi III dorsoplantarly deeper. Distinguished from Namaortyx sperrgebietensis Mourer-Chauviréet al., 2011, in that tarsometatarsus proportionally more elongated. Differentiated from Scopelortyx klinghardtensis Mourer-Chauviré et al., 2015, in that distal end of femur less excavated by sulcus intercondylaris, and proximal portion of condylus medialis more gradually merging with shaft. |
Measurements
No measurements are available
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| Source: o = order | |||||
| References: Livezey 1997, Marsh 1875 | |||||