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Shakiremys colombiana
Discussion
https://zoobank.org/NomenclaturalActs/46c7be26-cfa4-4843-baae-d98ab7a796db
Derivation of name. From Colombia, the country where the species was discovered.
Taxonomy
Shakiremys colombiana was named by Cadena et al. (2025). Its type specimen is VPPLT-1730, a partial skeleton (omplete skull (Figs 2, 3A−C, 4A−B) and articulated shell (carapace and plastron) (Figs 5, 6), belonging to a single individual (Fig. 1D).), and it is a 3D body fossil. Its type locality is Finca La Fortuna, which is in a Serravallian terrestrial horizon in the La Victoria Formation of Colombia. It is the type species of Shakiremys.
Synonymy list
| Year | Name and author |
|---|---|
| 2025 | Shakiremys colombiana Cadena et al. figs. 2, 3A−C, 4A−B, 5−7 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Shakiremys colombiana Cadena et al. 2025
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Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| E. Cadena et al. 2025 | Distinguished from other podocnemidids by the following combination of features: (1) an intermediate roofing of the temporal emargination, exhibiting a nearly straight posterior edge, less expanded than in, for example Peltocephalus dumerilianus, Erymnochelys madagascariensis and Dacquemis paleomorpha, and more expanded than in Podocnemis spp. Neochelys spp. and Caninemys tridentata; (2) a jugal retracted from the orbital margin, enabling contact between the maxilla and postorbital; (3) a very shallow cheek emargination enabling contact between the jugal and the quadrate as in Pe. dumerilianus and Er. madagascariensis (polymorphic); (4) an elongated postorbital bone that prevents contact between the parietal and jugal, different from Podocnemis spp.; (5) a narrow interorbital bar, as in Podocnemis spp., different from Pe. dumerilianus, Er. madagascariensis, Neochelys spp., Stupendemys geographica and stereogenyins; (6) orbits facing dorsolaterally, as in Podocnemis spp. and Ca. tridentata, differing from Pe. dumerilianus, Er. madagascariensis, St. geographica, Neochelys spp., Da. paleomorpha, and most stereogenyins; (7) an absent fossa precolumelaris, as in Po. expansa and Pe. dumerilianus; (8) an elongated triangular interparietal scute reaching nearly to the contact between the parietals and supraoccipital, as in Podocnemis spp. (except Po. vogli) and Er. madagascariensis, differing from the parallel lateral margins seen in Pe. dumerilianus, St. geographica and most stereogenyins; (9) foramen palatinum posterius located at the palatine–pterygoid suture as in Pe. dumerilianus and St. geographica; (10) shape of the condylus mandibularis much wider than long, with anterior and posterior edges straight to concave, making it shorter at midline, different from the ‘kidney bean’ of Podocnemis spp.; (11) processus paraoccipitalis of the opisthotic almost at the same level of the most posterior corner of the squamosal, with a small marginal notch between them, as can occur in Po. sextuberculata and Po. vogli; (12) aperture narium externa partially covered by the prefrontals forming a medial acute tip as in Podocnemis spp.; (13) at least one accessory ridge on the ventral surface of the premaxilla–maxilla suture, as in Podocnemis spp. (one or two ridges) and Ca. tridentata (two ridges); (14) a flatter skull, as in Podocnemis spp., contrasting with the higher, more globular skull of Pe. dumerilianus and Er. madagascariensis; (15) a flatter and more rectangular carapace outline, in contrast to the moderately domed and oval-elongated outline seen in all other podocnemidids; (16) an anterior carapace embayment, similar to St. geographica; (17) peripherals 2 with a short, protruding anteromedial margin, similar to St. geographica (female specimens) (Cadena et al. 2021); (18) a very large costal 1, twice the length of costal 2; (19) vertebral scute 1 narrower than the others and not reaching peripherals 1, as in juvenile specimens of St. geographica; (20) seven neural bones, which is the most common condition in Podocnemis spp. and Pe. dumerilianus, differing from Er. madagascariensis and St. geographica (juvenile specimen) (Cadena et al. 2021), which have only six; (21) gular scute reaching the pectorals and completely separating the humerals; and (22) anterior plastral lobe margin almost straight, shared only by some specimens of St. geographica within podocnemidids. |
Measurements
No measurements are available
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| Source: f = family, c = class, subp = subphylum | |||||
| References: Hendy et al. 2009, Uetz 2005, Carroll 1988, Kiessling 2004 | |||||
Age range: Serravallian or 13.82000 to 11.63000 Ma
Collections (2 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Serravallian | Colombia (Huila) | Shakiremys colombiana (type locality: 275773 275774) |