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Spermophilus howelli

Mammalia - Rodentia - Sciuridae

Taxonomy
Citellus howelli was named by Hibbard (1941). It is a 3D body fossil. Its type locality is Rexroad KU Locality 3, which is in a Blancan terrestrial horizon in the Rexroad Formation of Kansas.

It was recombined as Spermophilus howelli by Hemming (1956), Shotwell (1968), Kurten and Anderson (1980), Kelly (1994).

Sister species lacking formal opinion data
S. matachisensis

Synonymy list
YearName and author
1941Citellus howelli Hibbard
1956Spermophilus howelli Hemming
1963Citellus howelli Black
1968Spermophilus howelli Shotwell
1980Spermophilus howelli Kurten and Anderson p. 212
1994Spermophilus howelli Kelly

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
—Therapsida()
infraorderCynodontia()
RankNameAuthor
—Mammaliamorpha
—Mammaliaformes
classMammalia
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—EuarchontogliresMurphy et al. 2001
—GliriformesWyss and Meng 1996
—Glires()
—Simplicidentata()
orderRodentiaBowdich 1821
superfamilySciuroidea
familySciuridae
tribeMarmotini
genusSpermophilusCuvier 1825
specieshowelli()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Spermophilus howelli Hibbard 1941
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Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: semifossorialg
Diet: granivoref
Diet 2: frugivoref
Reproduction: viviparoussubc
Created: 2012-05-15 10:05:16
Modified: 2012-05-14 21:55:07
Source: g = genus, f = family, subc = subclass, c = class, subp = subphylum
References: Carroll 1988, Lillegraven 1979, Ji et al. 2002, Fernández et al. 2000, Nowak 1991, Hendy et al. 2009

Age range: Blancan or 4.70000 to 1.40000 Ma

Collections (5 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Hemphillian - Blancan9.4 - 1.4USA (Indiana) Spermophilus howelli (18320)
Blancan4.7 - 1.4USA (Texas) Spermophilus howelli (20023)
Blancan4.7 - 1.4USA (Nevada) Spermophilus howelli (19998)
Blancan4.7 - 1.4USA (Kansas) Spermophilus howelli (type locality: 19936)
Blancan4.7 - 1.4USA (Washington) Spermophilus howelli (20062)