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Emydinae

Osteichthyes - Testudines - Emydidae

Taxonomy
Emydinae was named by Rafinesque (1815). It is extant. Its type is Emys. It was considered monophyletic by Bickham (1975).

It was reranked as the tribe Emydinae by Perrier (1928).

It was assigned to Trionychidae by Lydekker (1889); to Testudinidae by Perrier (1928), Estes (1964), Estes and Tihen (1964), Kuhn (1966), Melentis (1966), Estes and Berberian (1970); and to Emydidae by Bickham (1975), Ernst and Barbour (1989), Holman (2002), van Dijk et al. (2014), Bourque (2016).

Synonymy list
YearName and author
1815Emydinae Rafinesque
1889Emydinae Lydekker p. 22
1928Emydinae Perrier p. 3113
1964Emydinae Estes p. 99
1964Emydinae Estes and Tihen p. 464
1966Emydinae Kuhn p. 26
1966Emydinae Melentis
1970Emydinae Estes and Berberian p. 5
1975Emydinae Bickham p. 201
1989Emydinae Ernst and Barbour p. 136
2002Emydinae Holman p. 439
2014Emydinae van Dijk et al.
2016Emydinae Bourque

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
RankNameAuthor
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
classReptilia
—Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
—Pantestudinoidea
superfamilyTestudinoidea
familyEmydidaeRafinesque 1815
subfamilyEmydinaeRafinesque 1815
subfamilyEmydinaeRafinesque 1815

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Emydinae Rafinesque 1815
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G. Actinemys Agassiz 1857
G. Clemmys Ritgen 1828
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†Clemmys gaudryi Depéret 1885
Clemmys guttata Schneider 1792
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Invalid names: Clemmys punctata Schoepff 1792 [synonym]
†Clemmys schansiensis Ping 1932
†Clemmys ukoi Bachmayer 1957
G. Emyda Geoffroy Saint-Hilaire 1809
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Emyda vittata Peters 1854
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Invalid names: Emyda ceylonensis Gray 1855 [synonym]
G. †Wilburemys Martin and Mallory 2011
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†Wilburemys yakimensis Martin and Mallory 2011
Diagnosis
ReferenceDiagnosis
E. Perrier 1928Espèces aquatiques (Tortues d'eau douce).
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: freshwaterf
Locomotion: actively mobilec
Life habit: aquaticf
Diet: omnivoref
Reproduction: oviparousf
Created: 2005-08-26 14:20:16
Modified: 2005-08-26 16:20:16
Source: f = family, c = class, subp = subphylum
References: Carroll 1988, Hendy et al. 2009, Uetz 2005, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Late/Upper Maastrichtian to the top of the Holocene or 72.20000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 66.0 Ma

Collections (46 total)


Time interval Ma Country or state Original ID and collection number
Lancian72.2 - 66.0USA (Wyoming) Emydinae indet. (14505 14519 14538 14566)
Late/Upper Maastrichtian72.2 - 66.0USA (Wyoming) Emydinae indet. (14585)
Late/Upper Maastrichtian72.2 - 66.0USA (Montana) Emydinae indet. (13142)
Wasatchian54.9 - 50.5Canada (Northwest Territories) Emydinae indet. (15583)
Oligocene33.9 - 23.04France (Auvergne) Clemmys jungi (36690)
Whitneyan31.8 - 29.5USA (Florida) Pseudemys sp. (17341)
Late/Upper Oligocene27.3 - 23.04Georgia Clemmys sp. (182744)
Late/Upper Oligocene - Early/Lower Miocene27.3 - 15.98Kazakhstan Clemmys kasachstanica (74901)
Early/Lower Miocene23.04 - 15.98Russian Federation Clemmys sp. (43647)
Barstovian16.3 - 12.5USA (Nebraska) Chrysemys sp. (18284) Clemmys sp. (18141 27507) Emydinae indet. (18102 18206 18208)
Tortonian11.63 - 7.246Austria (Lower Austria) Clemmys ukoi (217032)
Late/Upper Miocene11.63 - 5.333India (Gujarat) Emyda vittata (87489)
MN 911.1 - 9.67Slovakia (Trnava) Emydinae indet. (69719)
Hemphillian9.4 - 4.7Costa Rica (Puntarenas) Emydinae indet. (144596)
Hemphillian9.4 - 4.7USA (Washington) Wilburemys yakimensis (232055)
Zanclean5.333 - 3.6France (Pyrénées-Orientales) Emys gaudryi (178671 178672)
Pliocene5.333 - 2.58Ukraine (Odessa) Clemmys pidoplickai (76548)
Pliocene5.333 - 2.58China (Shansi) Clemmys schansiensis (195442)
Pliocene5.333 - 2.58Russian Federation (Rostovskaya Oblast') Clemmys indet. (39409)
Ruscinian5.09 - 3.55China (Shanxi) Clemmys sp. (42273)
MN 154.16 - 3.55Poland Clemmys sp. (33396)
Late/Upper Pliocene - Early/Lower Pleistocene3.6 - 0.774Pakistan Emyda vittata (92070)
MN 163.55 - 2.245Poland Clemmys sp. (33473)
Pleistocene2.58 - 0.0117USA (California) Clemmys sp. (237076)
Pleistocene2.58 - 0.0117Japan (Ryukyu Islands) Clemmys sp. (48721)
Irvingtonian1.4 - 0.21USA (California) Clemmys sp. (20173 20176 20177 20387 20431)
Middle Pleistocene0.774 - 0.129USA (California) Clemmys sp. (20639)
Late/Upper Pleistocene (interglacial)0.129 - 0.0117USA (California) Clemmys sp. (20646)
Late/Upper Pleistocene (late glacial)0.129 - 0.0117USA (Florida) Chrysemys sp. (74361)
Late/Upper Pleistocene (high glacial)0.129 - 0.0117USA (Texas) Pseudemys sp. (74470)
Late/Upper Pleistocene0.129 - 0.0117USA (California) Clemmys sp. (73752)
Late/Upper Pleistocene (glacial)0.129 - 0.0117USA (California) Clemmys sp. (155393)
Holocene0.0117 - 0.0Canada (Quebec) Clemmys guttata (224189)