PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Borhyaeninae

Osteichthyes - Sparassodonta

Taxonomy

Borhyaeninae was named by Ameghino (1894). Its type is Boryhyaena.

It was assigned to Borhyaenidae by Marshall (1977), Marshall (1978), Patterson and Marshall (1978), de Muizon (1999), Goin et al. (2007).

Subtaxa

Acrocyon, Arctodictis, Borhyaena (syn. Pseudoborhyaena, Conodonictis, Dynamictis), Eutemnodus (syn. Apera)

Synonyms

Sparassodontidae was named by Roger (1896).

It was synonymized subjectively with Borhyaeninae by Marshall (1978).
Conodonictidae was named by Ameghino (1935).

It was synonymized subjectively with Borhyaeninae by Marshall (1978).

Synonymy list

Year	Name and author
1894	Borhyaeninae Ameghino
1896	Sparassodontidae Roger p. 16
1935	Conodonictidae Ameghino p. 131
1977	Borhyaeninae Marshall p. 419
1978	Borhyaeninae Marshall
1978	Borhyaeninae Patterson and Marshall
1999	Borhyaeninae de Muizon p. 506
2007	Borhyaeninae Goin et al.

Is something missing? Join the Paleobiology Database and enter the data

Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
class	Osteichthyes
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota

Rank	Name	Author
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()
—	Mammaliamorpha
—	Mammaliaformes
class	Mammalia
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
subclass	Metatheria	()
—	Marsupialiformes
order	Sparassodonta	()
superfamily	Borhyaenoidea	Ameghino 1894
—	Borhyaenidae	()
subfamily	Borhyaeninae	Ameghino 1894
subfamily	Borhyaeninae	Ameghino 1894

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Borhyaeninae Ameghino 1894

show all | hide all

G. †Acrocyon Ameghino 1887

hide

†Acrocyon riggsi Sinclair 1930

†Acrocyon sectorius Ameghino 1887

Invalid names: Acrocyon eguianus Mercerat 1891 [synonym]

G. †Arctodictis Mercerat 1891

hide

†Arctodictis munizi Mercerat 1891

†Arctodictis sinclairi Marshall 1978

G. †Borhyaena Ameghino 1887

hide

†Borhyaena macrodonta Ameghino 1902

Invalid names: Pseudoborhyaena longaeva Ameghino 1902 [synonym]

†Borhyaena tuberata Ameghino 1887

Invalid names: Acrocyon patagonensis Mercerat 1891 [synonym], Arctodictis australis Mercerat 1891 [synonym], Borhyaena fera Ameghinor 1891 [synonym], Borhyaena sanguinaria Ameghino 1894 [synonym], Borhyaena zitteli Ameghino 1894 [synonym], Conodonictis exterminator Ameghino 1891 [synonym], Conodonictis rapax Cabrera 1927 [synonym], Conodonictis saeva Ameghino 1891 [synonym]

Invalid names: Conodonictis Ameghino 1891 [synonym], Dynamictis Ameghino 1891 [synonym], Pseudoborhyaena Ameghino 1902 [synonym]

G. †Eutemnodus Bravard 1858

hide

†Eutemnodus acutidens Riggs and Patterson

†Eutemnodus americanus Bravard 1858

Invalid names: Apera sanguinaria Ameghino 1886 [synonym], Hyaenodon sudamericanus Burmeister 1891 [synonym]

†Eutemnodus propampina Ameghino 1891

Invalid names: Apera Ameghino 1886 [synonym]

Invalid names: Conodonictidae Ameghino 1935 [synonym], Sparassodontidae Roger 1896 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Maximum body mass:

23.31 kgs

Minimum body mass:

23.31 kgs

Body mass estimate:

23.31 kg

Environment:

terrestrialsubo

Locomotion:

actively mobilec

Life habit:

scansorialsubc

Diet:

carnivoresubo

Reproduction:

viviparoussubc

Created:

2013-05-24 23:51:13

Modified:

2013-05-24 08:51:13

Source: s = species, subo = suborder, subc = subclass, c = class, subp = subphylum

References: Clemens 1979, Goin 1997, Lillegraven 1979, Carroll 1988, Vizcaíno et al. 2010, Hendy et al. 2009

Age range: base of the Lutetian to the top of the Laventan or 48.07000 to 11.00000 Ma

Collections (35 total)

Time interval	Ma	Country or state	Original ID and collection number
Lutetian	48.07 - 41.03	Argentina (Chubut)	B. gen. et sp. indet. C (133039)
Middle Eocene - Early/Lower Miocene	48.07 - 15.98	Argentina (Chubut)	Borhyaena riggsi (176461)
Deseadan	29.0 - 21.0	Brazil (Sao Paulo)	Borhyaeninae indet. (152418)
Deseadan	29.0 - 21.0	Bolivia	Proborhyaena sp. (95688)
Deseadan	29.0 - 21.0	Bolivia (La Paz)	Borhyaeninae indet. (133596)
Early/Lower Miocene	23.04 - 15.98	Argentina (Chubut)	Borhyaena macrodonta (236575)
Colhuehuapian	21.0 - 17.5	Argentina (Chubut)	Acrocyon riggsi (176459) Borhyaena macrodonta (176436) Borhyaeninae indet., Pseudoborhyaena macrodonta, Pseudoborhyaena longaeva, Acrocyon riggsi, Arctodictis sinclairi (141368)
Santacrucian	17.5 - 16.3	Argentina (Santa Cruz)	Acrocyon sectorius (176580 176581 176594) Arctodictis munizi (176603 206242) Arctodictis munizi, Borhyaena tuberata (142533) Borhyaena tuberata (142529 176565 176566 176570 176576 176578 176582 176585 176596 176602 176636 210638 210786) Borhyaena tuberata, Acrocyon sectorius (176593) Borhyaena tuberata, Arctodictis munizi (176563 176587) Borhyaena zitteli, Borhyaena tuberata, Arctodictis munizi (27313)
Friasian	16.3 - 15.5	Chile (Aisén)	Borhyaena tuberata (139845)
Colloncuran	15.5 - 13.8	Argentina (Neuquen)	Arctodictis sp. (28611)
Laventan	13.8 - 11.0	Colombia (Huila)	Arctodictis sp. (195258)