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Alioramus remotus
Taxonomy
Alioramus remotus was named by Kurzanov (1976). Its type specimen is PIN 3141/1, a partial skeleton, and it is a 3D body fossil. Its type locality is Nogon Tsav (PIN 3141) [SMPE/SMGE], which is in a Maastrichtian terrestrial siliciclastic in the Beds of Nogon Tsav Formation of Mongolia. It is the type species of Alioramus. It was considered monophyletic by Currie (2000).
Synonyms
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Synonymy list
Year | Name and author |
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1976 | Alioramus remotus Kurzanov p. 94 figs. 1-9 |
1977 | Alioramus remotus Perle p. 111 |
1986 | Alioramus remotus Gauthier p. 9 |
1988 | Alioramus remotus Paul p. 327 |
1990 | Alioramus remotus Molnar et al. p. 190 |
1991 | Alioramus remotus Molnar p. 166 |
1991 | Alioramus remotus Weishampel et al. p. 209 |
1992 | Alioramus remotus Currie p. 246 |
1997 | Alioramus remotus Barsbold p. 448 |
1997 | Alioramus remotus Osmólska p. 472 |
1997 | Alioramus remotus Vickaryous and Ryan p. 488 |
1999 | Alioramus remotus Knoll et al. p. 105 |
2000 | Alioramus remotus Currie p. 448 |
2001 | Alioramus remotus Holtz, Jr. pp. 68-69 fig. 7.2 |
2003 | Alioramus remotus Currie p. 654 |
2003 | Alioramus remotus Currie et al. p. 231 |
2003 | Alioramus remotus Rauhut p. 43 |
2004 | Alioramus remotus Holtz, Jr. p. 113 |
2009 | Alioramus altai Brusatte et al. p. 17261 figs. 1-3 |
2009 | Alioramus altai Brusatte and et al. |
2010 | Alioramus altai Carr p. 1220 |
2011 | Alioramus altai Carr et al. |
2011 | Alioramus remotus Carr et al. |
2011 | Alioramus altai Fowler et al. p. 3 |
2011 | Alioramus altai Hone et al. p. 495 |
2011 | Alioramus remotus Hone et al. p. 501 |
2012 | Alioramus altai Bell et al. p. 188 |
2012 | Alioramus remotus Bell et al. p. 188 |
2013 | Alioramus altai Dalman p. 241 |
2013 | Alioramus altai Poropat and Kear p. 105 |
2013 | Alioramus altai Thomson et al. p. 71 |
2013 | Alioramus remotus Thomson et al. p. 71 |
2014 | Alioramus altai Hendrickx and Mateus p. 29 |
2014 | Alioramus altai Lü et al. |
2014 | Alioramus remotus Lü et al. |
2017 | Alioramus remotus Carr et al. p. 9s |
2017 | Alioramus altai Carr et al. p. 16s |
2018 | Alioramus altai Frederickson et al. p. 6 |
2018 | Alioramus altai McDonald et al. p. 2 |
2018 | Alioramus remotus McDonald et al. p. 2 |
2019 | Alioramus altai Brougham et al. p. 13 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Alioramus remotus Kurzanov 1976
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Invalid names: Alioramus altai Brusatte 2009 [synonym]
Diagnosis
Reference | Diagnosis | |
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S. L. Brusatte et al. 2009 (Alioramus altai) | A. altai is a tyrannosaurid theropod possessing the following autapomorphies: an accessory pneumatic fenestra posterodorsal to promaxillary fenestra of maxilla; enlarged and elongated maxillary fenestra (length:depth ratio of 1.9); a laterally projecting horn on the jugal; a thick ridge on the dorsal surface of the ectopterygoid; a palatine pneumatic recess extending posteriorly beyond the posterior margin of the vomeropterygoid process; 20 dentary alveoli; an anteroposteriorly elongate anterior mylohyoid foramen of the splenial; a thin epipophysis on the atlantal neurapophysis that terminates at a sharp point; a pneumatic pocket on the anterior surface of the cervical transverse processes; an external pneumatic foramina on the dorsal ribs; and an anterodorsally inclined midline ridge on the lateral surface of the ilium. Many of these features are present on elements not preserved in the holotype of A. remotus (see SI Text).
A. altai is distinguished from the holotype of A. remotus, which is at approximately the same ontogenetic stage judging by the slight 3% difference in reconstructed skull length between the two specimens, by: s.c. flange on maxilla (the lateral surface of the maxilla extends dorsally to form a narrow slot between itself and the antorbital fossa below the ventral margin of the antorbital fenestra, which is absent in A. remotus); three less-developed rugosities on the nasal (as opposed to six more-prominent rugosities in A. remotus); an anterior process of quadratojugal terminates posterior to the anterior margin of the lateral temporal fenestra; a squamosal anterior process that extends anterior to the anterior margin of the lateral temporal fenestra; and an epipterygoid not bifurcated ventrally. A. altai also possesses three differences with A. remotus that are size-related in other tyrannosaurids but may be significant given the similar size of the holotypes: 17 maxillary and 20 dentary alveoli (16 and 18, respectively, in A. remotus); a single dorsoventral groove between the basal tubera (groove bifurcated by ridge in A. remotus); and a tapering anterior process of the parietals overlapping frontals on the midline (larger, rectangular process in A. remotus). Although representing a juvenile animal (see Histological and Ontogenetic Analysis), the holotype of A. altai can be distinguished from juveniles of the contemporary Tarbosaurus by numerous characters. Namely, Tarbosaurus subadults have a deeper maxilla, a deeper tooth-bearing region of the maxilla, fewer teeth anterior to the antorbital fossa, a rounder maxillary fenestra, more closely spaced maxillary and promaxillary fenestrae, low and indistinct lacrimal horns, low nasal rugosities, a larger postorbital horn, and a considerably lower tooth count in the maxilla and dentary (11, 12). |
Measurements
No measurements are available
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Source: subo = suborder, o = order | |||||
References: Marsh 1875, Benton 1983 |
Age range: Maastrichtian or 72.20000 to 66.00000 Ma
Collections (2 total)
Time interval | Ma | Country or state | Original ID and collection number |
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Maastrichtian | Mongolia (Bayankhongor) | Alioramus remotus (type locality: 91909) | |
Maastrichtian | Mongolia (Omnogov) | Alioramus altai (113009) |