PBDB Taxon

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Squalodontidae

Osteichthyes - Cetacea - Squalodontidae

Taxonomy

Squalodontidae was named by Brandt (1873). It is not extant. It was considered monophyletic by Uhen et al. (2008), Barnes and Reynolds (2009).

It was assigned to Odontoceta by Ameghino (1894); to Cetacea by Newton (1891), Rhoads (1903); to Odontocete by Hay (1902), Hay (1930); to Squaloceti by Abel (1914), Abel (1919), Dal Piaz (1929), Van Deinse (1931); to Archaeoceti by Paquier (1894), Benham (1935); to Squalodontoidea by Simpson (1945), Rothausen (1961), Rothausen (1968), Ginsburg and Janvier (1971), Barnes et al. (1985), Pilleri (1987), Vidal (1991), McKenna and Bell (1997), Dubrovo and Sanders (2000); to Platanistoidea by Muizon (1984), Muizon (1987), Muizon (1990), Muizon (1994), Fordyce and Barnes (1994), Fordyce et al. (1995), Rice (1998), Fordyce and de Muizon (2001), Rice (2002), Gingerich (2005), Godfrey et al. (2006), Barnes (2006), Bianucci and Landini (2007), Whitmore and Kaltenbach (2008), Uhen et al. (2008), Rice (2009), Barnes and Reynolds (2009), Bianucci et al. (2011), Carone and Marra (2014), Aguirre-Fernández and Fordyce (2014), Marx et al. (2016); to Stegoceti by Boessenecker and Geisler (2023); to Odontoceti by Lydekker (1887), Cope (1889), Cope (1890), Zittel (1894), Flores (1895), Trouessart (1898), Trouessart (1904), Case (1904), Abel (1905), True (1908), Jaekel (1911), Miller (1923), Zittel (1925), Kellogg (1928), Keyes (1973), Whitmore and Sanders (1977), Whitmore (1984), Pilleri (1985), Carroll (1988), Benton (1993), Sach and Heizmann (2001), Geisler and Sanders (2003), Cahuzac et al. (2005), Godfrey and Lambert (2023); and to Odontoceti by Winge (1921), Slijper (1936), Geisler et al. (2011), Berta (2017), Nelson et al. (2025), Nelson et al. (2025), Nelson et al. (2025).

Subtaxa

Austrosqualodon, Austrosqualodon trirhizodonta, Eosqualodon, Pachyodon, Portheodon, Squalodon (syn. Delphinoides, Macrophoca, Phocodon, Rhytisodon, Trirhizodon, Kelloggia, Arionius, Smilocomptus, Rhizoprion, Crenidelphinus), Squalodon andrewi, Squalodon atlanticus, Squalodon bordae, Squalodon catulli, Squalodon melitensis (syn. Squalodon scillae), Squalodontinae, Tangaroasaurus, Tangaroasaurus kakanuiensis, Zeuglodon vredense

Synonyms

Zeuglodon vredense was named by Landois (1844) [No type speciemen designated].

It was considered a nomen dubium by Stromer (1903); it was considered a nomen nudum by Kellogg (1936); it was recombined as Basilosaurus vredensis by Trouessart (1904), Kalmykov (2012).
Austrosqualodon trirhizodonta was named by Climo and Baker (1972). It is not extant. Its type specimen is DM 1627. Its type locality is Fossil Point, which is in a Duntroonian marine siltstone in the Abel Head Formation of New Zealand.

It was considered a nomen dubium by Fordyce and de Muizon (2001); it was considered a nomen dubium by Nelson et al. (2025).
Tangaroasaurus kakanuiensis was named by Benham (1935) [Otago University]. Its type specimen is OU C.03.18 GL426, a mandible, and it is a 3D body fossil. Its type locality is All Day Bay, which is in a Langhian marine claystone in the Gee Greensand Formation of New Zealand.

It was considered a nomen dubium by Nelson et al. (2025).
Squalodon andrewi was named by Benham (1942) [Otago University. The holotype tooth is missing, but the natural limestone cast of the tooth, an adjacent tooth, and part of the palate, is held as OU C.77.21, formerly part of collection C.75.25.]. Its type specimen is OU C.77.21, a set of teeth, and it is a 3D body fossil. Its type locality is Clarendon, which is in a Chattian carbonate limestone in the Milburn Limestone Formation of New Zealand.

It was considered an invalid subgroup of Squalodontoidea by Fordyce (1980); it was considered a nomen dubium by Fordyce (1991); it was considered an invalid subgroup of Squalodontidae by Fordyce and Roberts (2009).
Pachyodon was named by Meyer (1838). It is not extant.

It was considered a nomen dubium by Kellogg (1923); it was synonymized subjectively with Squalodon by Lydekker (1887), Zittel (1894), Trouessart (1898), Dal Piaz (1916), Zittel (1925).

It was assigned to Zeuglodontae by Bronn (1853).
Pachyodon catulli was named by Molin (1859) [No type specimen number is listed by Mollin (1859). Kellogg (1923) states that the type specimen was in the Museum of the University of Padova, Italy, but are now lost.]. It is a 3D body fossil. Its type locality is Libano, which is in an Aquitanian marine sandstone in the Libano Sandstone Formation of Italy. It is the type species of Trirhizodon.

It was recombined as Squalodon catulli by Brandt (1873), de Zigno (1876), Lydekker (1887), Roger (1887), Lioy (1896), Rothausen (1968), Sanders et al. (1982), Pilleri (1985), Bianucci and Landini (2002), Symeonidis et al. (2004); it was synonymized subjectively with Squalodon bariensis by Dal Piaz (1901); it was synonymized subjectively with Squalodon bariense by Dal Piaz (1916); it was recombined as Trirhizodon catulli by Kellogg (1923); it was misspelled as Squalodon catulloi by Zittel (1894), Trouessart (1904); it was considered a nomen dubium by Nelson et al. (2025).
Macrophoca atlantica was named by Leidy (1856) [ANSP 11218 was designated as the lectotype by Dooley, 2003 with ANSP 11217 and ANSP 11219 as paralectotypes]. Its type specimen is ANSP 11218, a set of teeth (3 molar teeth), and it is a 3D body fossil. Its type locality is Shiloh Marl, which is in a Burdigalian marginal marine marl in the Kirkwood Formation of New Jersey. It is the type species of Macrophoca.

It was recombined as Squalodon atlantica by Cope (1867), Roger (1887), Case (1904), Kellogg (1923); it was recombined as Squalodon atlanticum by Dal Piaz (1916); it was recombined as Basilosaurus atlantica by Allen (1916); it was considered an invalid subgroup of Squalodontidae by Dooley (2003); it was recombined as Squalodon atlanticus by Cope (1868), Cope (1868), Leidy (1869), Cope (1875), Lydekker (1887), Cope (1890), Zittel (1894), Trouessart (1898), Hay (1902), Rhoads (1903), Trouessart (1904), Hay (1930), Uhen et al. (2008).
Delphinus bordae was named by Gervais (1852) [Kellogg (1923) states that the type specimen was formerly in the collection of H. Arnozan, "directeur de l'Asile departmental des alienes de la Charente-Inferieur, France.]. It is a 3D body fossil. Its type locality is Leognan, which is in a Burdigalian marine sandstone in France.

It was recombined as Champsodelphis bordae by Gervais (1852), Pictet (1853), Scott (1873); it was synonymized subjectively with Squalodon grateloupii by Trouessart (1898); it was recombined as Squalodon bordae by Abel (1905), Kellogg (1923); it was considered a nomen dubium by Dal Piaz (1916), Nelson et al. (2025).
Phoca melitensis was named by Blainville (1840) [On p 46 this species is called Phoca dubia melitensis Kellogg (1923) states that the type specimen is in the Cambridge University Museum, England.]. It is a 3D body fossil. Its type locality is Malta General, which is in a Chattian/Langhian marine phosphorite in the Globigerina Limestone Formation of Malta.

It was recombined as Squalodon melitensis by McCoy (1867), Brandt (1873), Kellogg (1923), Rothausen (1968), Symeonidis et al. (2004); it was synonymized subjectively with Squalodon scillae by Trouessart (1898); it was considered a nomen dubium by Nelson et al. (2025).
Portheodon was named by Gill (1872) [Gill assigns this genus to Cope, but no earlier reference to the name can be found.].

It was synonymized subjectively with Squalodon by Zittel (1894); it was considered a nomen nudum by Palmer (1904).

It was assigned to Cynorcidae by Gill (1872).
Mesoceti was named by Dames (1894). It is not extant.

It was synonymized subjectively with Squalodontidae by Trouessart (1898).
Phocodon scillae was named by Agassiz (1841) [No type specimen number listed by Agassiz (1841). No type specimen number listed by Agassiz (1841).]. Its type specimen is Sedgwick Museum, Cambridge, a mandible (mandible fragment with three teeth), and it is a 3D body fossil. Its type locality is Wied l-Isquof, which is in a Burdigalian/Langhian marine phosphorite in the Upper Main Phosphorite Conglomerate Formation of Malta.

It was recombined as Squalodon scillae by Brandt (1873), Brandt (1874), Trouessart (1898), Trouessart (1904), Kellogg (1922); it was synonymized subjectively with Squalodon melitensis by Kellogg (1923), Rothausen (1968), Bianucci et al. (2011).

Synonymy list

Year	Name and author
1838	Pachyodon Meyer p. 414
1840	Phoca melitensis Blainville pp. 46, 51 figs. plate 10, fig. 4
1841	Phocodon scillae Agassiz p. 236
1844	Zeuglodon vredense Landois p. 49
1852	Champsodelphis bordae Gervais p. 153
1852	Delphinus bordae Gervais p. 153 figs. pl. 41, Fig. 8
1853	Pachyodon Bronn p. 774
1853	Champsodelphis bordae Pictet p. 383
1859	Pachyodon catulli Molin p. 126 figs. Plates 1-2
1867	Squalodon atlantica Cope
1867	Squalodon melitensis McCoy p. 145
1868	Squalodon atlanticus Cope p. 166
1869	Squalodon atlanticus Leidy p. 416
1872	Portheodon Gill
1873	Squalodon catulli Brandt
1873	Squalodon melitensis Brandt p. 330
1873	Squalodon scillae Brandt p. 330
1873	Squalodontidae Brandt p. 576
1873	Champsodelphis bordae Scott p. 67
1874	Squalodon scillae Brandt p. III
1875	Squalodon atlanticus Cope p. 363
1876	Squalodon catulli de Zigno p. 24
1887	Squalodon atlanticus Lydekker p. 75
1887	Squalodon catulli Lydekker p. 75
1887	Squalodontidae Lydekker p. 75
1887	Squalodon atlantica Roger p. 27
1887	Squalodon catulli Roger p. 27
1889	Squalodontidae Cope p. 876
1890	Squalodontidae Cope p. 602
1890	Squalodon atlanticus Cope p. 603
1891	Squalodontidae Newton p. 75
1894	Squalodontidae Ameghino p. 436
1894	Mesoceti Dames
1894	Squalodontidae Paquier p. 5
1894	Zeuglodon vredense Zittel p. 168
1894	Squalodontidae Zittel p. 169
1894	Squalodon catulloi Zittel p. 170
1894	Squalodon atlanticus Zittel p. 171
1895	Squalodontidae Flores p. 11
1896	Squalodon catulli Lioy p. 765
1898	Zeuglodon vredensis Trouessart p. 1010
1898	Squalodontidae Trouessart p. 1012
1898	Squalodon scillae Trouessart p. 1013
1898	Squalodon atlanticus Trouessart p. 1015
1902	Squalodontidae Hay p. 588
1902	Squalodon atlanticus Hay p. 589
1903	Squalodon atlanticus Rhoads p. 247
1903	Squalodontidae Rhoads p. 247
1904	Squalodon atlantica Case p. 6
1904	Squalodontidae Case p. 6
1904	Basilosaurus vredensis Trouessart p. 753
1904	Squalodontidae Trouessart p. 754
1904	Squalodon catulloi Trouessart p. 755
1904	Squalodon scillae Trouessart p. 755
1904	Squalodon atlanticus Trouessart p. 756
1905	Squalodon bordae Abel p. 27
1905	Squalodontidae Abel p. 46
1908	Squalodontidae True p. 391
1911	Squalodontidae Jaekel p. 231
1914	Squalodontidae Abel p. 221
1916	Basilosaurus atlantica Allen
1916	Squalodon atlanticum Dal Piaz p. 8
1916	Phocodon scillae Dal Piaz p. 14
1919	Squalodontidae Abel p. 762
1921	Squalodontidae Winge p. 7
1922	Squalodon scillae Kellogg p. 126
1923	Squalodon atlantica Kellogg p. 7
1923	Squalodon bordae Kellogg p. 9
1923	Trirhizodon catulli Kellogg p. 11
1923	Squalodon melitensis Kellogg p. 24
1923	Squalodontidae Miller p. 40
1925	Squalodontidae Zittel p. 85
1928	Squalodontidae Kellogg p. 32 figs. Table 1
1929	Squalodontidae Dal Piaz p. 73
1930	Squalodontidae Hay p. 580
1930	Squalodon atlanticus Hay p. 581
1931	Squalodontidae Van Deinse p. 33
1935	Tangaroasaurus kakanuiensis Benham p. 235
1935	Squalodontidae Benham p. 243
1936	Squalodontidae Slijper p. 543
1942	Squalodon andrewi Benham pp. 267-268
1945	Squalodontidae Simpson p. 100
1961	Squalodontidae Rothausen p. 113
1968	Squalodontidae Rothausen p. 87
1968	Squalodon catulli Rothausen p. 91 figs. Table 1
1968	Squalodon melitensis Rothausen p. 91 figs. Table 1
1971	Squalodontidae Ginsburg and Janvier p. 167
1972	Austrosqualodon trirhizodonta Climo and Baker p. 67 figs. Fig. 2
1973	Austrosqualodon trirhizodonta Keyes
1973	Tangaroasaurus kakanuiensis Keyes p. 381
1973	Squalodon andrewi Keyes p. 389
1973	Squalodontidae Keyes p. 389
1977	Squalodontidae Whitmore and Sanders p. 305
1979	Tangaroasaurus kakanuiensis Fordyce p. 740
1980	Austrosqualodon trirhizodonta Fordyce p. iii figs. Table 2
1980	Tangaroasaurus kakanuiensis Fordyce p. iii figs. Table 2
1982	Austrosqualodon trirhizodonta Fordyce p. 423
1982	Squalodon catulli Sanders et al. p. H120
1984	Squalodontidae Muizon p. 68
1984	Squalodontidae Whitmore p. 230
1985	Squalodontidae Barnes et al. p. 25
1985	Squalodon catulli Pilleri p. 11
1985	Squalodontidae Pilleri p. 11
1987	Squalodontidae Muizon p. 13
1987	Squalodontidae Pilleri p. 147
1988	Squalodontidae Carroll
1990	Squalodontidae Muizon p. 312
1991	Austrosqualodon trirhizodonta Fordyce p. 1182
1991	Tangaroasaurus kakanuiensis Fordyce p. 1182
1991	Squalodontidae Vidal p. 5
1993	Squalodontidae Benton p. 761
1994	Squalodontidae Fordyce and Barnes p. 429 figs. Table 1
1994	Squalodontidae Muizon p. 136
1995	Squalodontidae Fordyce et al. p. 379
1997	Squalodontidae McKenna and Bell p. 373
1998	Squalodontidae Rice
2000	Squalodontidae Dubrovo and Sanders p. 580
2001	Squalodontidae Fordyce and de Muizon p. 178
2001	Squalodontidae Sach and Heizmann p. 42
2002	Squalodon catulli Bianucci and Landini p. 21
2002	Squalodontidae Rice p. 231 figs. Table 1
2003	Squalodontidae Geisler and Sanders p. 28
2004	Squalodon catulli Symeonidis et al. p. 319 figs. Table 2
2004	Squalodon melitensis Symeonidis et al. p. 319 figs. Table 2
2005	Squalodontidae Cahuzac et al. p. 420
2005	Squalodontidae Gingerich p. 237 figs. Table 15.1
2006	Squalodontidae Barnes p. 32
2006	Squalodontidae Godfrey et al. p. 68A
2007	Squalodontidae Bianucci and Landini p. 45 figs. Table 2.1
2008	Squalodon atlanticus Uhen et al. p. 583
2008	Squalodontidae Uhen et al. p. 583
2008	Squalodontidae Whitmore and Kaltenbach p. 183
2009	Squalodontidae Barnes and Reynolds p. 487
2009	Austrosqualodon trirhizodonta Fordyce and Roberts p. 553
2009	Tangaroasaurus kakanuiensis Fordyce and Roberts p. 553
2009	Squalodontidae Rice p. 235 figs. Table 1
2011	Squalodontidae Bianucci et al. p. 560
2011	Squalodontidae Geisler et al. p. 5 figs. Table 1
2012	Basilosaurus vredensis Kalmykov p. 179
2014	Squalodontidae Aguirre-Fernández and Fordyce p. 195
2014	Squalodontidae Carone and Marra p. 122
2016	Squalodontidae Marx et al. p. 120
2017	Squalodontidae Berta p. 160
2023	Squalodontidae Boessenecker and Geisler p. 152
2023	Squalodontidae Godfrey and Lambert p. 53
2025	Squalodontidae Nelson et al. p. 3

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
class	Osteichthyes
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()
—	Mammaliamorpha

Rank	Name	Author
—	Mammaliaformes
class	Mammalia
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Boreoeutheria
—	Laurasiatheria
—	Scrotifera
—	Euungulata
—	Artiodactylamorpha
—	Artiodactyla	()
—	Whippomorpha
order	Cetacea
—	Pelagiceti
—	Neoceti
suborder	Odontoceti
family	Squalodontidae
family	Squalodontidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Squalodontidae Brandt 1873

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G. †Austrosqualodon Climo and Baker 1972

G. †Eosqualodon Rothausen 1968

†Eosqualodon langewieschei Rothausen 1968

†Eosqualodon latirostris Capellini 1903

G. †Squalodon Grateloup 1840

†Squalodon barbarus Mchedlidze and Aslanova 1968

†Squalodon bellunensis Dal Piaz 1901

†Squalodon calvertensis Kellogg 1923

†Squalodon grateloupii von Meyer 1843

Invalid names: Delphinoides gratelupi Pedroni 1845 [objective synonym], Squalodon bariensis Jourdan 1861 [synonym], Squalodon burgueti Gervais 1849 [synonym], Squalodon grateloupii Gervais 1846 [objective synonym], Squalodon kelloggi Rothausen 1968 [synonym], Squalodon zitteli Paquier 1894 [synonym]

†Squalodon murdochi Godfrey and Lambert 2023

†Squalodon peregrinus Dal Piaz 1971

†Squalodon servatus Meyer 1841

†Squalodon whitmorei Dooley 2005

Invalid names: Arionius Meyer 1841 [synonym], Crenidelphinus Laurillard 1844 [synonym], Delphinoides Pedroni 1845 [synonym], Kelloggia Mchedlidze 1976 [synonym], Macrophoca Leidy 1856 [synonym], Phoca pedronii Gervais 1849 [invalid subgroup], Phocodon Agassiz 1841 [synonym], Priscodelphinus validus Du Bus 1872 [nomen nudum], Rhizoprion Jourdan 1861 [synonym], Rhytisodon Costa 1852 [synonym], Smilocomptus Gervais 1849 [synonym], Squalodon antverpiensis Van Beneden 1861 [nomen dubium], Squalodon antverpiensis antverpiensis Rothausen 1968 [nomen dubium], Squalodon quaternarium Forsyth Major 1879 [nomen nudum], Trirhizodon Cope 1890 [synonym]

G. †Tangaroasaurus Benham 1935

Invalid names: Austrosqualodon trirhizodonta Climo and Baker 1972 [nomen dubium], Mesoceti Dames 1894 [synonym], Pachyodon Meyer 1838 [nomen dubium], Portheodon Gill 1872 [nomen nudum], Squalodon andrewi Benham 1942 [invalid subgroup], Squalodon atlanticus Leidy 1856 [invalid subgroup], Squalodon bordae Gervais 1852 [nomen dubium], Squalodon catulli Molin 1859 [nomen dubium], Squalodon melitensis Blainville 1840 [nomen dubium], Squalodontinae Rothausen 1968 [empty], Tangaroasaurus kakanuiensis Benham 1935 [nomen dubium], Zeuglodon vredense Landois 1844 [nomen nudum]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

hydroxyapatitesubo

Form:

roller-shapedo

Ontogeny:

modification of partso

Environment:

marine, freshwatersubo

Locomotion:

actively mobileo

Life habit:

aquatico

Depth habitat:

surfaceo

Diet:

carnivoresubo

Reproduction:

viviparoussubo

Created:

2005-03-06 14:21:39

Modified:

2005-09-22 15:42:08

Source: subo = suborder, o = order

Reference: Uhen 2004

Age range: base of the Duntroonian to the top of the Tortonian or 27.30000 to 7.24600 Ma

Collections (99 total)

Time interval	Ma	Country or state	Original ID and collection number
Duntroonian	27.3 - 25.2	New Zealand (N. W. Nelson)	Austrosqualodon trirhizodonta (13100)
Chattian	27.3 - 23.04	Germany	Squalodon sp. (57119)
Chattian	27.3 - 23.04	Germany (Mecklenburg-Vorpommern)	Squalodontidae indet. (80366)
Chattian	27.3 - 23.04	USA (South Carolina)	Squalodon sp. (97550 206943) Squalodon sp., S. Genus X, S. Genus Y, S. Genus Z (112635) Squalodontidae indet. (70957)
Chattian	27.3 - 23.04	Mexico (Baja California Sur)	Squalodontidae indet. (66426 186570)
Chattian	27.3 - 23.04	Azerbaijan	Squalodon barbarus (45737)
Chattian	27.3 - 23.04	Australia (Victoria)	Parasqualodon wilkinsoni (46166)
Chattian	27.3 - 23.04	New Zealand (South Otago)	Lophocephalus parki (45654) Squalodontidae indet., Squalodon andrewi (45653)
Chattian	27.3 - 23.04	Japan (Hokkaido)	Squalodontidae indet. (45786)
Chattian	27.3 - 23.04	Japan	Squalodon sp. (51103 51105)
Chattian	27.3 - 23.04	USA (California)	Squalodontidae indet. (11601)
Chattian	27.3 - 23.04	Japan (Fukuoka)	Squalodontidae indet. (45834)
Chattian	27.3 - 23.04	Germany (Nordrhein-Westfalen)	Eosqualodon langewieschei (37598)
Late/Upper Oligocene - Early/Lower Miocene	27.3 - 15.98	New Zealand (North Otago)	Squalodontidae indet. (87888)
Chattian - Langhian	27.3 - 13.82	Malta	Phocodon scillae (48661)
Waitakian	25.2 - 21.7	New Zealand (South Canterbury)	Squalodon andrewi (45655 45656)
Waitakian	25.2 - 21.7	New Zealand	Squalodontidae indet. (74755)
Aquitanian	23.04 - 20.45	Italy	Pachyodon catulli, Squalodon bariensis, Squalodon bellunensis, Squalodon peregrinus (47623) Squalodon bellunensis (59850)
Aquitanian	23.04 - 20.45	Austria	Squalodon bellunensis (76129)
Aquitanian	23.04 - 20.45	France	Squalodon sp. (70242)
Aquitanian	23.04 - 20.45	USA (Virginia)	Squalodon calvertensis (70517)
Aquitanian	23.04 - 20.45	Italy (Vicenza)	Squalodon bariensis (49395)
Aquitanian - Burdigalian	23.04 - 15.98	USA (New Jersey)	Squalodon sp. (50944)
Aquitanian - Burdigalian	23.04 - 15.98	Malta	Squalodon sp. (122120)
Early/Lower Miocene	23.04 - 15.98	Slovakia	Squalodon sp. (47461)
Early/Lower Miocene	23.04 - 15.98	Germany	Squalodontidae indet. (70360)
Early/Lower Miocene	23.04 - 15.98	Italy	Squalodontidae indet. (116545)
Early/Lower Miocene	23.04 - 15.98	Greece (Macedonia)	Squalodon bellunensis (47083)
Aquitanian - Langhian	23.04 - 13.82	USA (North Carolina)	Squalodon sp., Squalodontidae indet., Squalodon calvertensis (48887)
Aquitanian - Serravallian	23.04 - 11.63	USA (Maryland)	Squalodon calvertensis (231816)
Early/Lower Miocene - Middle Miocene	23.04 - 11.63	Australia (Victoria)	Squalodontidae indet. (51330)
Miocene	23.04 - 5.333	France (Côtes-d'Armor)	Squalodon sp. (148402)
Miocene	23.04 - 5.333	France	Squalodon gervaisii, Squalodon grateloupii (93327)
Miocene	23.04 - 5.333	Germany (Westphalia)	Zeuglodon vredense (73868)
Miocene	23.04 - 5.333	United Kingdom (England)	Squalodon antverpiensis (153047)
Miocene	23.04 - 5.333	Netherlands	Phoca ambigua (37599)
Burdigalian	20.45 - 15.98	Germany	Arionius servatus (47425) Squalodon zitteli (48663)
Burdigalian	20.45 - 15.98	USA (Delaware)	Squalodon calvertensis (18579)
Burdigalian	20.45 - 15.98	USA (California)	Squalodontidae indet. (55691)
Burdigalian	20.45 - 15.98	USA (Virginia)	Squalodon whitmorei (57951 57953 57959)
Burdigalian	20.45 - 15.98	Germany (Baden-Württemberg)	Squalodontidae indet. (49102)
Burdigalian	20.45 - 15.98	Italy	Squalodon bariensis, Squalodon bellunense (47187) Squalodon sp. (70814 207869) Squalodontidae indet. (47427)
Burdigalian	20.45 - 15.98	USA (New Jersey)	Macrophoca atlantica (18587)
Burdigalian	20.45 - 15.98	Switzerland	Squalodontidae indet. (69539)
Burdigalian	20.45 - 15.98	Venezuela (Falcon)	Squalodontidae indet. (43063)
Burdigalian	20.45 - 15.98	Australia (South Australia)	Squalodontidae indet. (51407)
Burdigalian	20.45 - 15.98	France (Drôme)	Rhizoprion bariensis (48662)
Burdigalian	20.45 - 15.98	Switzerland (Zurich)	Squalodon sp. (46523)
Burdigalian	20.45 - 15.98	USA (Maryland)	Squalodon calvertensis (45493) Squalodon calvertensis, Squalodon whitmorei (71810) Squalodon whitmorei (57960)
Burdigalian	20.45 - 15.98	Italy (Sicily)	Squalodon bariensis (49396)
Burdigalian	20.45 - 15.98	Slovakia	Squalodon ehrlichii (48642)
Burdigalian	20.45 - 15.98	France (Bordeaux)	Squalodon grateloupii, Squalodon bordae, Delphinoides gratelupi, Phoca pedronii (48660)
Burdigalian	20.45 - 15.98	France	Squalodon grateloupi (57732) Squalodon sp. (52061)
Burdigalian	20.45 - 15.98	Switzerland (Aargau)	Squalodon sp. (46524)
Burdigalian	20.45 - 15.98	Argentina (Chubut)	Squalodontidae indet. (45886)
Burdigalian - Langhian	20.45 - 13.82	Belgium	Squalodon sp. (114830)
Burdigalian - Messinian	20.45 - 5.333	Italy (Apulia)	Squalodon sp. (48657) Squalodon sp., Squalodon antverpiensis, Squalodon scillae, Phocodon scillae (47125)
Early/Lower Barstovian	16.3 - 12.5	USA (Florida)	Squalodon sp. (18549)
Langhian	15.98 - 13.82	USA (Maryland)	Squalodon murdochi (45506 60379) Squalodon sp. (55517 70819)
Langhian	15.98 - 13.82	USA (Virginia)	Squalodon calvertensis (79793)
Langhian	15.98 - 13.82	Malta	Squalodon sp. (81668)
Langhian	15.98 - 13.82	Italy (Sicillia)	Squalodon bariensis (28617)
Langhian	15.98 - 13.82	New Zealand (North Otago)	Tangaroasaurus kakanuiensis (45659)
Langhian - Serravallian	15.98 - 11.63	Belgium (Antwerpen)	Squalodon antverpiensis, Priscodelphinus validus (51680)
Langhian - Serravallian	15.98 - 11.63	Belgium	Squalodon sp. A (59123)
Middle Miocene	15.98 - 11.63	Costa Rica (Limon)	Squalodon sp. (124711)
Middle Miocene - Pleistocene	15.98 - 0.0117	USA (Massachusetts)	Squalodon atlanticus (77694)
Badenian	13.82 - 12.8	Belgium	Squalodon sp. (95167)
Serravallian	13.82 - 11.63	Japan (Ishikawi)	Squalodon sp. (76038)
Serravallian	13.82 - 11.63	Italy	Squalodontidae indet., Squalodon bellunensis (48656)
Serravallian	13.82 - 11.63	France (Gironde)	Smilocomptus burgueti (85340)
Serravallian	13.82 - 11.63	USA (Virginia)	Squalodon calvertensis (71801) Squalodon sp. (57113 187324)
Serravallian	13.82 - 11.63	USA (Maryland)	Squalodon atlanticus (57782)
Serravallian	13.82 - 11.63	France	Squalodon sp. (85338)
Tortonian	11.63 - 7.246	Italy	Squalodon sp. (80394)
Late/Upper Miocene - Early/Lower Pliocene	11.63 - 3.6	Chile	Delphinus sp. (73840)