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Gryposaurus notabilis

Reptilia - Hadrosauridae

Taxonomy
Gryposaurus notabilis was named by Lambe (1914). Its type specimen is NMC 2278, a partial skeleton, and it is a 3D body fossil. Its type locality is RTMP Quarry 35, Red Deer River, which is in a Campanian channel sandstone in the Dinosaur Park Formation of Canada.

It was recombined as Kritosaurus notabilis by Brown (1914), Sinclair and Granger (1914), Gilmore (1924), Huene (1927), Hay (1930), Russell (1930), Gilmore (1933), Wilfarth (1940), Lull and Wright (1942), Vialli (1960), Kuhn (1964), Tatarinov (1964), Russell (1964), Pinna (1967), Brambilla (1972) and Baird and Horner (1979); it was recombined as Hadrosaurus notabilis by Horner (1979) and Weishampel and Weishampel (1983); it was synonymized subjectively with Kritosaurus navajovius by Hunt and Lucas (1992).

Synonyms
Synonymy list
YearName and author
1914Kritosaurus notabilis Brown p. 379
1914Gryposaurus notabilis Lambe p. 145
1914Kritosaurus notabilis Sinclair and Granger
1915Gryposaurus notabilis Lambe p. 117
1916Gryposaurus notabilis Lambe p. 195
1919Kritosaurus incurvimanus Parks
1919Gryposaurus notabilis Parks p. 51
1919Kritosaurus incurvimanus Parks p. 51 figs. 1-13
1920Kritosaurus incurvimanus Parks pp. 5-9
1920Gryposaurus notabilis Parks pp. 7-8
1924Kritosaurus incurvimanus Gilmore p. 22
1924Kritosaurus notabilis Gilmore p. 34
1924Kritosaurus incurvimanus Parks p. 382
1927Kritosaurus incurvimanus Huene p. 260
1927Kritosaurus notabilis Huene p. 260
1930Kritosaurus incurvimanus Hay p. 213
1930Kritosaurus notabilis Hay p. 214
1930Kritosaurus incurvimanus Russell p. 148
1930Kritosaurus notabilis Russell p. 148
1933Kritosaurus notabilis Gilmore p. 62
1935Kritosaurus incurvimanus Parks p. 193
1936Kritosaurus incurvimanus Nagao p. 186
1939Kritosaurus incurvimanus Nagao p. 144
1940Kritosaurus notabilis Wilfarth p. 10
1942Kritosaurus notabilis Lull and Wright p. 167 fig. 59
1942Kritosaurus incurvimanus Lull and Wright pp. 169-170
1960Kritosaurus notabilis Vialli p. 173
1960Kritosaurus incurvimanus Vialli p. 176
1964Kritosaurus incurvimanus Kuhn p. 21
1964Kritosaurus notabilis Kuhn p. 21
1964Kritosaurus incurvimanus Russell p. 12
1964Kritosaurus notabilis Russell p. 12
1964Kritosaurus notabilis Tatarinov p. 563 fig. 612
1966Gryposaurus notabilis Russell p. 17
1966Kritosaurus incurvimanus Russell p. 27
1967Kritosaurus notabilis Pinna p. 145
1972Kritosaurus notabilis Brambilla pp. 316-317
1979Kritosaurus incurvimanus Baird and Horner p. 19
1979Kritosaurus notabilis Baird and Horner p. 19
1979Hadrosaurus notabilis Horner
1983Hadrosaurus notabilis Weishampel and Weishampel p. 44
1987Kritosaurus incurvimanus Powell p. 150
1990Gryposaurus notabilis Weishampel and Horner p. 556
1991Gryposaurus notabilis Weishampel et al. p. 202 fig. 8
1992Gryposaurus incurvimanus Horner p. 13
1992Gryposaurus notabilis Horner p. 15
1993Gryposaurus notabilis Hunt and Lucas p. 80
1998Gryposaurus notabilis Godefroit et al. p. 26
1998Kritosaurus incurvimanus Godefroit et al. p. 47
2001Gryposaurus notabilis Eberth et al. p. 58
2001Kritosaurus incurvimanus Eberth et al. p. 58
2001Gryposaurus incurvimanus Horner et al. p. 7
2001Gryposaurus notabilis Horner et al. p. 7
2004Gryposaurus incurvimanus Horner et al. p. 440
2004Gryposaurus notabilis Horner et al. p. 440
2005Gryposaurus incurvimanus Currie p. 5
2005Gryposaurus notabilis Currie p. 5
2005Gryposaurus incurvimanus Ryan and Evans p. 336
2005Gryposaurus notabilis Ryan and Evans p. 336
2006Gryposaurus incurvimanus Kirkland et al. p. 311 fig. 13
2006Gryposaurus notabilis Kirkland et al. p. 311 fig. 13
2006Gryposaurus incurvimanus Lund and Gates p. 272
2006Gryposaurus notabilis Lund and Gates p. 272
2006Gryposaurus incurvimanus Prieto-Márquez et al. p. 88
2006Gryposaurus notabilis Prieto-Márquez et al. p. 88
2006Gryposaurus notabilis Serrano-Brañas et al. p. 75
2007Gryposaurus incurvimanus Gates and Sampson p. 353
2007Gryposaurus notabilis Gates and Sampson p. 353
2007Gryposaurus incurvimanus Gates et al. p. 925
2008Gryposaurus incurvimanus Longrich p. 994
2008Gryposaurus notabilis Longrich p. 994
2009Gryposaurus notabilis Evans et al. p. 1316
2010Gryposaurus incurvimanus Gates et al. p. 168
2010Gryposaurus notabilis Gates et al. p. 168
2010Gryposaurus notabilis Prieto-Marquez p. 850
2010Gryposaurus notabilis Prieto-Marquez and Salinas p. 835 fig. 20
2010Gryposaurus notabilis Prieto-Márquez p. 5 fig. 2
2010Gryposaurus notabilis Prieto-Márquez and Norell p. 6
2010Gryposaurus notabilis Prieto-Márquez and Wagner p. 1243 fig. 3
2011Gryposaurus notabilis Eberth and Evans p. 18
2011Gryposaurus notabilis Gates et al. p. 800
2011Gryposaurus notabilis Prieto-Márquez p. 63
2012Gryposaurus notabilis Bell p. 2
2012Gryposaurus notabilis Godefroit et al. p. 551
2012Gryposaurus notabilis Godefroit et al. p. 4
2012Gryposaurus incurvimanus Godefroit et al. p. 13
2012Gryposaurus notabilis Prieto-Márquez p. 512
2012Gryposaurus notabilis Ramírez-Velasco et al. p. 392
2012Gryposaurus notabilis Saveliev et al. p. 80
2013Gryposaurus notabilis Campione et al. p. 66
2013Gryposaurus notabilis Cruzado-Caballero et al.
2013Gryposaurus notabilis Prieto-Marquez
2014Gryposaurus notabilis Bell p. 575
2014Gryposaurus notabilis Bell et al. p. 177
2014Gryposaurus notabilis Campione p. 208
2014Gryposaurus notabilis Farke and Herrero p. 191
2014Gryposaurus notabilis Gates et al. p. 156
2014Gryposaurus notabilis Prieto-Márquez p. 296
2014Gryposaurus notabilis Prieto-Márquez and Wagner p. 595
2014Gryposaurus notabilis Ramírez-Velasco et al. p. 350
2014Gryposaurus incurvimanus Rivera-Sylva and Carpenter p. 160
2014Gryposaurus notabilis Tsogtbaatar et al. p. 115
2014Gryposaurus notabilis Xing et al. p. 1819
2015Gryposaurus notabilis Tan et al. p. 249
2016Gryposaurus notabilis Lehman et al. p. 345
2017Gryposaurus notabilis Bertozzo et al. p. 12
2017Gryposaurus incurvimanus Cruzado-Caballero and Powell p. 11
2017Gryposaurus notabilis Cruzado-Caballero and Powell p. 11
2018Gryposaurus notabilis Fondevilla et al. p. 8
2018Gryposaurus incurvimanus Fondevilla et al. p. 21
2019Gryposaurus notabilis Drysdale et al. p. 15
2019Gryposaurus notabilis Takasaki et al. p. 2
2020Gryposaurus notabilis Dalla Vecchia p. 612
2021Gryposaurus notabilis Kobayashi et al. p. 5
2021Gryposaurus notabilis McDonald et al. p. 13

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
classReptilia
subclassEureptilia()
—Romeriida
—Diapsida()
—Archosauromorpha(Huene 1946)
—Crocopoda
RankNameAuthor
—ArchosauriformesGauthier 1986
—Eucrocopoda
—Archosauria()
informalAvemetatarsalia
—Ornithodira
—Dinosauromorpha
—Dinosauriformes
—Dinosauria()
—Ornithischia()
—Genasauria
—Cerapoda
—Ornithopoda()
—Clypeodonta
—Iguanodontia()
—Dryomorpha
—Styracosterna
—Hadrosauriformes
—Hadrosauroidea()
familyHadrosauridae
subfamilyHadrosaurinaeLambe 1918
tribeKritosauriniLapparent and Lavocat 1955
genusGryposaurus
speciesnotabilis

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Gryposaurus notabilis Lambe 1914
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Invalid names: Gryposaurus incurvimanus Parks 1919 [synonym]
Diagnosis
ReferenceDiagnosis
T. A. Gates and S. D. Sampson 2007Hadrosaurids of the genus Gryposaurus with the following unique combination of characters: sigmoidal lateroventral margin of the maxilla; large nasal arch located near the orbits and rising above the level of the frontals; narrow ‘U’-shaped posterior margin of the external naris; squamosal posterodorsal margin well above the level of the skull roof; infratemporal fenestra larger than the orbit; and rounded mandibular foramen. The skull of G. notabilis is generally more robust than that of G. latidens and G. incurvimanus, but less so than that of G. monumentensis.
T. A. Gates and S. D. Sampson 2007 (Gryposaurus incurvimanus)Hadrosaurids of the genus Gryposaurus possessing: dorsal premaxillary process that is more concave posteriorly than in other species of the genus; nasal arch smaller than in other species and positioned well anterior to the orbit and rising to a level below the frontals; jugal with a small spur on the posterior margin of the posteroventral flange; and slightly excavated ventral surface of the nasal hump. In addition, G. incurvimanus possesses the following unique combination of characters: nearly straight lateroventral margin of the maxilla; infratemporal fenestra larger than the orbit; and broad ‘U’-shaped posterior margin of the external naris. Gryposaurus incurvimanus is significantly more gracile than either G. notabilis or G. monumentensis. It has a smaller premaxillary lip than the other species, as well as premaxillae that are angled more steeply than in other species. Gryposaurus incurvimanus also possesses an anteroposteriorly shortened skull relative to the maximum skull depth.
A. Prieto-Marquez 2010Hadrosaurid dinosaur of the genus Gryposaurus characterized by the following autapomorphies: predentary denticles triangular and pointed, with bases that are wider or as wide as the denticles are tall; and infratemporal fenestra being both anteroposteriorly wider and dorsoventrally longer than the orbit. This species is also characterized by combining a nasal protuberance (that rises anterior to the orbits and above the level of the frontals) with a narrow U-shaped posterior margin of the narial fenestra, posterodorsal margin of the squamosal located well above the level of the skull roof, and rounded mandibular foramen (combination of characters modified from Gates and Sampson, 2007).
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yeso
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: gregariouso
Environment: terrestrialo
Locomotion: actively mobileo
Life habit: ground dwellingo
Diet: herbivoresubo
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2009-01-03 20:40:05
Modified: 2009-01-03 22:40:05
Source: subo = suborder, o = order
References: Benton 1983, Marsh 1875

Age range: Late/Upper Campanian or 83.50000 to 70.60000 Ma

Collections (10 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Campanian83.5 - 70.6Canada (Alberta) Gryposaurus notabilis (type locality: 58947) Gryposaurus sp. (59012 59013) Hadrosaurus sp. (59102) Kritosaurus notabilis (11919 55317 58946) Kritosaurus sp. (59119 96534)
Middle Campanian - Late/Upper Campanian83.5 - 70.6Canada (Alberta) Kritosaurus incurvimanus (46331)