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Macroolithus rugustus

Osteichthyes - Elongatoolithidae

Taxonomy
Oolithes rugustus was named by Young (1965). It is considered to be a form taxon. Its type specimen is IVPP V2788, an egg, and it is a 3D body fossil. Its type locality is IVPP Loc. 6224, Lashuyuan, which is in a Maastrichtian terrestrial sandstone in the Pingling Formation of China. It is the type species of Macroolithus.

It was recombined as Macroolithus rugustus by Zhao (1975), Zhao (1978), Zhao (1979), Zhao et al. (1991), Mikhailov (1994), Zhao (1994), Mikhailov (1997), Mikhailov (2000), Amo Sanjuán et al. (2000), Cousin (2002), Garcia et al. (2006), Wang et al. (2012), Wang et al. (2016), He et al. (2019), Simon et al. (2019), Tanaka et al. (2023).

Synonymy list
YearName and author
1965Oolithes rugustus Young p. 106
1973Oolithes rugustus Cheng et al. p. 23
1975Macroolithus rugustus Zhao p. 106 figs. Pl. 1-2:1
1978Macroolithus rugustus Zhao p. 214
1979Macroolithus rugustus Zhao p. 336
1980Oolithes rugustus Chao p. 61
1989Oolithes rugustus Hirsch p. 93
1991Macroolithus rugustus Zhao et al. p. 5
1992Oolithes rugustus Shen and Mateer p. 75
1994Macroolithus rugustus Mikhailov p. 117
1994Macroolithus rugustus Zhao p. 187
1997Macroolithus rugustus Mikhailov p. 64
2000Macroolithus rugustus Amo Sanjuán et al. p. 13
2000Macroolithus rugustus Mikhailov p. 569
2002Macroolithus rugustus Cousin p. 44
2006Macroolithus rugustus Garcia et al. p. 100 fig. 3
2012Macroolithus rugustus Wang et al. p. 407
2016Macroolithus rugustus Wang et al. p. 2
2019Macroolithus rugustus He et al. p. 353
2019Macroolithus rugustus Simon et al. p. 7
2023Macroolithus rugustus Tanaka et al. p. 202

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
RankNameAuthor
subclassEureptilia()
Romeriida
Diapsida()
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Saurischia()
Theropoda()
familyElongatoolithidae
genusMacroolithus
speciesrugustus()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Macroolithus rugustus Young 1965
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Diagnosis
ReferenceDiagnosis
K. E. Mikhailov 1994Predominant range of shell thicknesses (in equatorial part of egg) 1.1-1.3 mm, total range 0.8-1.5 mm (with elements of sculpture up to 1.9 mm, rarely up to 2.2 mm).
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yeso
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: solitaryo
Environment: terrestrialo
Locomotion: actively mobileo
Life habit: ground dwellingo
Diet: carnivoreo
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2004-12-14 12:50:08
Modified: 2004-12-14 14:50:08
Source: o = order
References: Kiessling 2004, Marsh 1875

Age range: base of the Campanian to the top of the Maastrichtian or 83.60000 to 66.00000 Ma

Collections (19 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Cretaceous100.5 - 66.0China (Jiangxi) Oolithes rugustus (71868)
Campanian83.6 - 72.2China (Guangdong) Macroolithus rugustus (275484 275485 275493)
Maastrichtian72.2 - 66.0Mongolia (Ovorhangai) Hadrosauridae indet. (13347)
Maastrichtian72.2 - 66.0China (Guangdong) Dinosauria indet. (71852) Macroolithus rugustus (24752 68149 68396 68397 68398 68399) Oolithes rugustus (71853 71856 71862)
Maastrichtian72.2 - 66.0Kazakhstan (Aqmola) Dinosauria indet. (13349)
Maastrichtian72.2 - 66.0China (Jiangxi) Macroolithus rugustus (224481)
Maastrichtian72.2 - 66.0Mongolia (Omnogov) Macroolithus rugustus (65335 165580)