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Hemicytheridae
Taxonomy
Hemicytherinae was named by Puri (1953). Its type is Hemicythere.
It was reranked as the family Hemicytheridae by Hazel and Holden (1971), Bertels (1973), Bertels (1975), Blaszyk (1987), Ayress (1995) and Liebau (2005).
It was assigned to Cytheridae by Bhatia and Mandwal (1960); to Trachyleberididae by Hazel (1983) and Carreño and Cronin (1993); to Cytheracea by Hazel and Holden (1971), Bertels (1973), Bertels (1975), Blaszyk (1987) and Ayress (1995); and to Trachyleberidoidea by Liebau (2005).
It was reranked as the family Hemicytheridae by Hazel and Holden (1971), Bertels (1973), Bertels (1975), Blaszyk (1987), Ayress (1995) and Liebau (2005).
It was assigned to Cytheridae by Bhatia and Mandwal (1960); to Trachyleberididae by Hazel (1983) and Carreño and Cronin (1993); to Cytheracea by Hazel and Holden (1971), Bertels (1973), Bertels (1975), Blaszyk (1987) and Ayress (1995); and to Trachyleberidoidea by Liebau (2005).
Subtaxa
Absonocytheropteron, Ambostracon, Aurila, Aurilini, Bradleyini, Campylocytherinae, Coquimbini, Echinocythereidinae, Elofsonella, Hemicythere (type genus), Hemicytheria, Heterocythereis, Mutilus, Neobuntonia, Nephokirkos, Oertliellinae, Orioninae, Orioninini, Patagonacythere, Pokornyellini, Procythereis, Quadracythere, Reussicythere, Robustaurila, Thaerocytherinae, Urocythereidinae
Synonymy list
Year | Name and author |
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1953 | Hemicytherinae Puri |
1960 | Hemicytherinae Bhatia and Mandwal p. 282 |
1971 | Hemicytheridae Hazel and Holden p. 2 |
1973 | Hemicytheridae Bertels |
1975 | Hemicytheridae Bertels p. 112 |
1983 | Hemicytherinae Hazel p. 99 |
1987 | Hemicytheridae Blaszyk p. 70 |
1993 | Hemicytherinae Carreño and Cronin p. 146 |
1995 | Hemicytheridae Ayress p. 916 |
2005 | Hemicytheridae Liebau |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. †Hemicytheridae Puri 1953
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Tr. †Aurilini Puri 1974
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Subfm. †Campylocytherinae Puri 1960
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Tr. †Leguminocythereidini Howe 1961
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G. Bensonocythere Hazel 1967
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†Bensonocythere blackwelderi Hazel 1983
†Bensonocythere bradyi Hazel 1983
†Bensonocythere gouldensis Hazel 1983
†Bensonocythere ricespitensis Hazel 1983
†Bensonocythere rugosa Hazel 1983
G. †Triginglymus
Tr. †Coquimbini Ohmert 1968
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G. Muellerina Bassiouni 1965
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†Muellerina bassiounii Hazel 1983
†Muellerina blowi Hazel 1983
†Muellerina ohmerti Hazel 1983
†Muellerina wardi Hazel 1983
Subfm. †Echinocythereidinae Hazel 1967
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G. Echinocythereis Puri 1953
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Echinocythereis echinata Sars 1866
†Echinocythereis leecreekensis Hazel 1983
Echinocythereis margaritifera Brady 1870
G. Elofsonella
G. †Hemicytheria
G. Mutilus
G. Neobuntonia Hartmann 1981
G. †Nephokirkos
Subfm. Orioninae Puri 1973
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Tr. †Orioninini Puri 1974
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G. Patagonacythere Hartmann 1962
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†Patagonacythere parvitenuis Hornibrook 1953
†Patagonacythere waihaoensis Ayress 1995
Tr. †Pokornyellini Puri 1974
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G. †Pokornyella Oertil 1956
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†Pokornyella devians Bonaduce et al. 1986
†Pokornyella italica Ruggieri et al. 1977
G. †Procythereis
G. Quadracythere Hornibrook 1952
G. †Reussicythere van den Bold 1966
G. Robustaurila Yajima 1982
Subfm. †Thaerocytherinae Hazel 1967
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G. Hermanites Puri 1955
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†Hermanites ascitus Hazel 1983
†Hermanites batequensis Carreño and Cronin 1993
†Hermanites huantraicoensis Bertels 1969
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†Jugosocythereis bermejoensis Carreño and Cronin 1993
†Jugosocythereis monrealensis Carreño and Cronin 1993
G. Poseidonamicus Benson 1972
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Poseidonamicus dinglei Boomer 1999
†Poseidonamicus major Benson 1972
Poseidonamicus minor Benson 1972
Poseidonamicus miocenicus Benson and Peypouquet 1983
†Poseidonamicus norrisi Yamaguchi et al. 2017
Poseidonamicus nudus Benson 1972
Poseidonamicus panopsus Whatley and Dingle 1989
Poseidonamicus pintoi Benson 1972
Poseidonamicus pseudorobustus Coles and Whatley 1989
Poseidonamicus riograndensis Benson and Peypouquet 1983
Poseidonamicus whatleyi Dingle 2003
Subfm. Urocythereidinae Hartmann and Puri 1974
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Invalid names: Oertliellinae Liebau 1975 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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Source: o = order, c = class | |||||
References: Jell and Adrain 2002, Kiessling 2004 |
Age range
Maximum range based only on fossils: base of the Early/Lower Albian to the top of the Holocene or 112.03000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 93.9 Ma
Minimum age of oldest fossil (stem group age): 93.9 Ma
Collections (181 total)
Time interval | Ma | Country or state | Original ID and collection number |
---|---|---|---|
Early/Lower Albian - Cenomanian | Argentina (Santa Cruz) | Hermanites sp. (234986) | |
Late/Upper Campanian | Saudi Arabia | Hermanites sp. (184843) | |
Maastrichtian | Brazil | Aurila sp. (210200) | |
Maastrichtian | Netherlands | Anticythereis euglypha (165094) | |
Late/Upper Maastrichtian | Netherlands | Anticythereis euglypha (32178 32179) | |
Late/Upper Maastrichtian - Danian | Argentina (Neuquén) | Occultocythereis sp. (232029) | |
Danian | Indian Ocean | Muellerina abyssicola (201904) | |
Danian | Argentina (Neuquén) | Hermanites huantraicoensis (232031) | |
Paleocene | USA (Missouri) | Hermanites gibsoni (232027) Hermanites gibsoni, Hermanites hadropleura (232028) Hermanites midwayensis (232025) Hermanites plusculmensis (232057) | |
Paleocene | Trinidad and Tobago | Hermanites collei (94013 94014 94021) Hermanites collei, Hermanites grimsdalei (94012 94018) | |
Early/Lower Eocene | Belgium (Vlaanderen) | Echinocythereis reticulatissima (94039) | |
Lutetian | India | Hermanites wadiai (145445) | |
Lutetian | Pakistan (Sindh) | Occultocythereis peristicia, Occultocythereis subspinellosa (94121) Quadracythere sp. 1, Hornibrookella arcana (94125) | |
Middle Eocene | Mexico (Baja California Sur) | Occultocythereis gioargaezi, Hermanites batequensis, Jugosocythereis bermejoensis (201740) Pokornyella sp., Jugosocythereis monrealensis (201739) | |
Middle Eocene | Hungary | Echinocythereis dadayana, Pokornyella inaequapunctata (171632 171633) | |
Bartonian | Bangladesh | Hornibrookella arcana (209194) Occultocythereis sp. A, Hornibrookella arcana (209195 209199) | |
Bartonian | USA (North Carolina) | Triginglymus sp. (60452) | |
Bartonian | USA (Alabama) | Triginglymus hyperochus (94040) | |
Priabonian | Tonga | Jugosocythereis sp., Nephokirkos sp., Pokornyella sp., Tongacythere kondoi (43006) | |
Priabonian | USA (Alabama) | Absonocytheropteron watervalleyensis, Echinocythereis jacksonensis, Hermanites dohmi, Hermanites hysonensis, Triginglymus gnythophoreus (94153) Absonocytheropteron watervalleyensis, Hermanites dohmi, Hermanites hysonensis, Triginglymus gnythophoreus (94157) Absonocytheropteron watervalleyensis, Hermanites hysonensis, Triginglymus gnythophoreus (94156) Echinocythereis jacksonensis, Hermanites dohmi, Hermanites hysonensis, Triginglymus gnythophoreus (94155) Echinocythereis jacksonensis, Hermanites dohmi, Triginglymus gnythophoreus (94154) | |
Priabonian | Libya | Hermanites sp., Occultocythereis sp. (77237) | |
Late/Upper Eocene | USA (Louisiana/Texas) | Echinocythereis jacksonensis (45017) | |
Late/Upper Eocene | New Zealand (Canterbury) | Hemicythere munida, Hemicythere gravis, Hermanites briggsi, Patagonacythere tricostata, Patagonacythere waihaoensis, Patagonacythere parvitenuis (161903) | |
Early/Lower Oligocene | Puerto Rico | Aurila socra, Jugosocythereis spinosa (94188 94190) Hemicythere gordoni, Jugosocythereis spinosa (94191) Jugosocythereis spinosa (94187 94189) | |
Early/Lower Oligocene | Mexico (Veracruz) | Echinocythereis mcguirti, Hemicythere sp., Hermanites sp., Jugosocythereis vicksburgensis (94173) Jugosocythereis vicksburgensis (94166) | |
Rupelian | Germany | Pokornyella limbata (95898) | |
Oligocene | Italy (Calabria) | Hermanites sp., Pokornyella amygdaliformis, Quadracythere sp. (9117) Hermanites sp., Pokornyella sp. (9116) | |
Oligocene | Antarctica | Quadracythere gazdzickii, Reussicythere piaseckii, Neobuntonia southshetlandensis (94115 94116) Reussicythere piaseckii (94114) | |
Late/Upper Oligocene | Puerto Rico | Aurila amygdala, Caudites sp., Hemicythere gordoni, Hermanites tschoppi, Jugosocythereis vicksburgensis, Jugosocythereis spinosa, Procythereis laresensis, Quadracythere brachypygaia, Quadracythere sp. (94194) Aurila amygdala, Hermanites tschoppi, Jugosocythereis vicksburgensis, Jugosocythereis laresensis (94197) Aurila amygdala, Hermanites tschoppi, Jugosocythereis vicksburgensis, Jugosocythereis laresensis, Procythereis laresensis, Quadracythere brachypygaia, Quadracythere sp. (94196) Hemicythere gordoni, Hermanites tschoppi, Jugosocythereis vicksburgensis, Jugosocythereis laresensis, Occultocythereis sp., Procythereis laresensis, Quadracythere brachypygaia (94198) Hemicythere gordoni, Jugosocythereis vicksburgensis, Quadracythere brachypygaia (94192) Jugosocythereis spinosa, Procythereis laresensis, Quadracythere sp. (94193) Jugosocythereis vicksburgensis, Jugosocythereis laresensis, Procythereis laresensis (94195) | |
Late/Upper Oligocene | USA (Louisiana) | Echinocythereis mcguirti, Jugosocythereis sp. (44521) | |
Late/Upper Oligocene | Antigua and Barbuda | Aurila socra (94182 94186) Aurila socra, Hermanites paijenborchiana, Hermanites sp., Jugosocythereis vicksburgensis (94185) Aurila socra, Quadracythere sp. (94177 94180) Hermanites paijenborchiana (94181) Jugosocythereis vicksburgensis (94183) Occultocythereis sp. (94179) Procythereis sp. (94184) | |
Chattian | USA (North Carolina) | Hermanites sp. (86499) | |
Aquitanian | South Africa (North Cape) | Ambostracon sp. (37451) | |
Early/Lower Miocene | Cuba (Sancti Spiritus) | Caudites sp., Procythereis deformis, Aurila cicatricosa (31183) | |
Early/Lower Miocene | Trinidad and Tobago | Echinocythereis asperrima (93910) | |
Early/Lower Miocene | Puerto Rico | Aurila galerita, Hermanites hutchisoni, Procythereis deformis, Quadracythere antillea (94200) Aurila galerita, Procythereis deformis (94202 94203) Aurila galerita, Procythereis deformis, Quadracythere antillea (94201) Jugosocythereis vicksburgensis, Procythereis deformis, Quadracythere antillea (94204) Procythereis deformis (94199) | |
Burdigalian | Italy (Sardinia) | Aurila bullapunctata (93939) Aurila bullapunctata, Aurila spp. (93937 93940) Aurila cicatricosa (93936) Aurila convexa, Pokornyella deformis (93944) Aurila spp. (93938) Aurila spp., Aurila convexa, Aurila opaca (93945) Aurila spp., Aurila convexa, Pokornyella deformis (93943) Pokornyella deformis (93942) | |
Burdigalian | India (Gujarat) | Hermanites haidingeri, Aurilla cicatricosa (93899) | |
Burdigalian | France | Aurila cauditoiles, Aurila vauclusi, Aurila ventroinflata (66288) | |
Burdigalian | South Africa (North Cape) | Ambostracon sp. (37487) | |
Burdigalian - Langhian | Turkey | Aurila soummamensis (206197 206230 206231) | |
Langhian | South Africa (North Cape) | Ambostracon sp., Hermanites sp. (37494) | |
Middle Miocene | Croatia | Aurila kollmanni, Aurila notata (68213) | |
Middle Miocene | Poland | Hermanites haidingeri, Quadracythere sp. (70539) | |
Serravallian | Poland | Aurila cicatricosa (138841 138844 138845 138846 138850 138851 138853 138855 138857 138858 138859 138860) Aurila larieyensis, Aurila cicatricosa (138852) Aurila larieyensis, Occultocythereis bituberculata, Aurila cicatricosa (138847) Occultocythereis bituberculata, Aurila cicatricosa (138842 138848 138849) Pokornyella deformis, Aurila cicatricosa (138843) | |
Sarmatian | Austria | Hemicytheria emphalodes, Hemicytheria omphalodes, Mutilus notata (73299) | |
Tortonian | Belgium (Antwerpen) | Hermanites excancellata, Muellerina latimarginata, Thaerocythere sp. (48092) | |
Tortonian | Panama (Colón) | Caudites medialis (93909) | |
Tortonian | Greece | Aurila albicans (138424) Aurila albicans, Aurila cicatricosa (138337 138429 138576 138578 138580 138581) Aurila albicans, Aurila longa, Aurila cicatricosa (138331 138579) Aurila albicans, Aurila longa, Aurila convexa, Aurila sp. B, Aurila cicatricosa (138336) Aurila albicans, Aurila longa, Aurila punctata, Aurila cicatricosa (138425) Aurila albicans, Aurila longa, Aurila punctata, Aurila sp. B, Aurila sp. 1, Aurila sp. 2, Aurila cicatricosa (138584) Aurila albicans, Aurila longa, Aurila punctata, Caudites calceolatus, Aurila cicatricosa (138312) Aurila albicans, Aurila sp. B, Aurila cicatricosa (138582) Aurila albicans, Pokornyella devians, Aurila cicatricosa (138428) Aurila albicans, Pokornyella italica (138431) Aurila albicans, Pokornyella italica, Aurila cicatricosa (138335) Aurila cicatricosa (138575) Aurila freudenthali, Aurila sp. 3, Aurila cicatricosa (138333) Aurila longa, Aurila cicatricosa (138443 138583) Aurila longa, Pokornyella italica, Aurila cicatricosa (138332) Caudites calceolatus, Pokornyella italica, Aurila cicatricosa (138334) | |
Late/Upper Miocene | Puerto Rico | Procythereis deformis, Quadracythere antillea (94205) Quadracythere antillea (94206) | |
Late/Upper Miocene | Colombia (Atlántico) | Caudites medialis (93908) Caudites medialis, Mutilus confragosus (93901 93904) Caudites nipeensis (93907) Caudites nipeensis, Caudites medialis, Mutilus confragosus (93900 93905 93906) Mutilus confragosus (93902 93903) | |
Late/Upper Miocene | Croatia | Hemicytheria brunnensis (68221) Hemicytheria brunnensis, Hemicytheria croatica, Hemicytheria major (68266) | |
Late/Upper Miocene | USA (Maryland) | Hemicythere conradi (191407) | |
Late/Upper Miocene | USA (Florida) | Hemicythere sp. (94805) | |
Messinian | Morocco | Mutilus sp. (34723 34724) | |
Messinian | Italy | Aurila sp., Pokornyella sp. (69899) | |
Messinian | Algeria | Mutilus sp., Urocythereis sp. (70879) | |
Messinian | Italy (Campania) | Aurila sp. (78633) | |
Messinian - Zanclean | North Atlantic | Echinocythereis whatleyi, Poseidonamicus sp., Poseidonamicus major (201874) Poseidonamicus major (201875) Poseidonamicus multipunctata, Poseidonamicus major (201900) | |
Early/Lower Pliocene | Greece (Rhodes) | Aurila convexa, Aurila sp., Urocythereis sp. (122994) | |
Early/Lower Pliocene | New Zealand | Hemicythere foveolata, Quadracythere biruga, Quadracythere sp. (73307) | |
Zanclean | USA (North Carolina) | Aurila laevicula, Bensonocythere calverti, Bensonocythere trapezoidalis, Bensonocythere whitei, Bensonocythere m, Bensonocythere oo, Bensonocythere pp, Bensonocythere qq, Bensonocythere rr, Echinocythereis planibasalis, Muellerina petersburgensis, Muellerina p, Proteoconcha gigantica, Proteoconcha mimica, Proteoconcha multipunctata, Proteoconcha tuberculata, Proteoconcha z, Thaerocythere schmidtae, Malzella evexa, Echinocythereis leecreekensis, Caudites paraasymmetricus, Muellerina ohmerti, Muellerina bassiounii, Muellerina wardi, Muellerina blowi, Thaerocythere carolinensis, Hermanites ascitus, Proteoconcha jamesensis, Bensonocythere bradyi, Bensonocythere blackwelderi, Bensonocythere gouldensis, Bensonocythere ricespitensis, Bensonocythere rugosa, Malzella conradi (52582) | |
Gelasian | Italy (Umbria) | Aurila convexa (200419) | |
Early/Lower Pleistocene | Panama | Caudites nipeensis (91912) | |
Early/Lower Pleistocene | USA (Florida) | Aurila conradi (41501) | |
Early/Lower Pleistocene | Italy | Aurila convexa, Aurila cruciata (176387) Aurila convexa, Aurila cruciata, Urocythereis sororcula, Urocythereis margaritifera (176385) | |
Calabrian | Italy (Umbria) | Aurila convexa (149603) Echinocythereis pustulata (200420) | |
Calabrian | USA (Florida) | Aurila conradi (41705) | |
Middle Pleistocene | USA (California) | Ambostracon costatum, Urocythereis californica (217054) | |
Late/Upper Pleistocene | India (Gujarat) | Caudites sp. (80599) | |
Holocene | France | Aurila convexa (56803) Aurila sp. (56806) | |
Holocene | India | Caudites sp. (76319) |