PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Gobisaurus domoculus

Reptilia - Ankylosauridae

Taxonomy

Gobisaurus domoculus was named by Vickaryous et al. (2001). Its type specimen is IVPP V12563, a skeleton (skull and unknown postcranial material), and it is a 3D body fossil. Its type locality is Maortu, which is in a Turonian terrestrial horizon in the Ulansuhai Formation of China. It is the type species of Gobisaurus. It was considered monophyletic by Vickaryous et al. (2004).

Synonyms

Zhongyuansaurus luoyangensis was named by Xu et al. (2007). Its type specimen is 41HIII-0002, a partial skeleton (nearly complete skull, fragmentary lower jaw, one cervical neural spine, one complete and five partial dorsal vertebrae, seven proximal caudal vertebrae, three ), and it is a 3D body fossil. Its type locality is Liufugou, Liudan, which is in an Aptian/Albian terrestrial horizon in the Haoling Formation of China.

It was recombined as Zhongyuanosaurus luoyangensis by Lü et al. (2007); it was misspelled as Zhongyuanosaurus luoyangensis by Lü (2008), Amiot et al. (2010), Jiang et al. (2011), Xu et al. (2011), Yang et al. (2013), Penkalski (2014) and Han et al. (2014); it was synonymized subjectively with Gobisaurus domoculus by Arbour and Currie (2016).

Synonymy list

Year	Name and author
2001	Gobisaurus domoculus Vickaryous et al. p. 1768 figs. 2-6
2004	Gobisaurus domoculus Vickaryous et al. p. 364
2007	Gobisaurus domoculus Lü et al. p. 888
2007	Zhongyuanosaurus luoyangensis Lü et al. p. 349
2007	Zhongyuansaurus luoyangensis Xu et al.
2008	Zhongyuanosaurus luoyangensis Lü p. 51
2010	Gobisaurus domoculus Amiot et al. p. 353
2010	Zhongyuansaurus luoyangensis Amiot et al. p. 353
2011	Zhongyuansaurus luoyangensis Jiang et al. p. 1150
2011	Zhongyuansaurus luoyangensis Xu et al. p. 213
2013	Gobisaurus domoculus Kirkland et al. p. 5
2013	Zhongyuansaurus luoyangensis Yang et al. p. 275
2014	Gobisaurus domoculus Han et al. p. 14 fig. 11
2014	Zhongyuanosaurus lauyangensis Han et al. p. 14 fig. 11
2014	Zhongyuansaurus luoyangensis Penkalski p. 618
2016	Gobisaurus domoculus Arbour and Currie p. 404
2017	Gobisaurus domoculus Arbour and Mallon p. 771

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()

Rank	Name	Author
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Ornithischia	()
—	Genasauria
—	Thyreophora	()
—	Ankylosauria	()
family	Ankylosauridae
genus	Gobisaurus
species	domoculus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Gobisaurus domoculus Vickaryous et al. 2001

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Invalid names: Zhongyuangosaurus luoyangensis Xu et al. 2007 [synonym]

Diagnosis

Reference	Diagnosis
M. K. Vickaryous et al. 2001	Large ankylosaurid with orbits 20% of cranial length and external nares 23% of cranial length; robust basipterygoid processes not fused to main pterygoid body; elongate premaxillary processes of vomers visible in palatal view. Similar to Shamosaurus in having delta-shaped dorsal profile, with premaxillae forming narrow apex. Differs from Shamosaurus in that cranium longer than wide; lacks discernible cranial sculpturing over antorbital area; maxillary tooth row length compared to overall cranium length relatively shorter; maximum premaxillary rostrum width greater than distance between the posteriormost maxillary teeth; reduced supraorbital bosses; and anterior surface of pterygoid vertically oriented.
V. M. Arbour and P. J. Currie 2016	Differs from all other ankylosaurs in the following combination of characters: rugose, amorphous frontonasal ornamentation not subdivided into distinct caputegulae; longitudinal groove on dorsal surface of premaxillary beak (also in Shamosaurus); squamosal horns shorter and more rounded than in Crichtonpelta; quadratojugal horn has centrally located apex, unlike Crichtonpelta; no prominences on nuchal crest, unlike Crichtonpelta; tooth row shorter relative to overall skull length than in Shamosaurus (26.7% in Gobisaurus, 40% in Shamosaurus); anterior surface of pterygoid is vertical, whereas it slopes anteroventrally in Shamosaurus.

Measurements

part	N	mean
femur diameter	1	80.0
fibual diameter	1	30.0
tibia diameter	1	70.0

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

gregariouso

Environment:

terrestrialo

Locomotion:

actively mobileo

Life habit:

ground dwellingo

Diet:

herbivoreo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2004-12-14 12:50:52

Modified:

2004-12-14 14:50:52

Source: o = order

Reference: Marsh 1875

Age range: base of the Aptian to the top of the Turonian or 125.00000 to 89.80000 Ma

Collections (2 total)

Time interval	Ma	Country or state	Original ID and collection number
Aptian - Albian	125.0 - 100.5	China (Henan)	Zhongyuansaurus luoyangensis (79687)
Turonian	93.9 - 89.8	China (Nei Mongol)	Gobisaurus domoculus (type locality: 66924)