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Protoceratops andrewsi

Osteichthyes - Protoceratopsidae

Taxonomy
Protoceratops andrewsi was named by Granger and Gregory (1923). Its type specimen is AMNH 6251, a skeleton, and it is a 3D body fossil. Its type locality is Shabarakh Usu (AMNH loc. 12049), which is in a Campanian eolian sandstone in the Djadokhta Formation of Mongolia.

Entered
by M. Carrano on 2003-02-06; modified by M. Carrano on 2013-10-27

Synonyms
Synonymy list
YearName and author
1923Protoceratops andrewsi Granger and Gregory pp. 1-9 figs. 32-34, 36
1923Protoceratops andrewsi T'an p. 123
1924Protoceratops andrewsi Joleaud p. 46
1924Protoceratops andrewsi Lull p. 245ff
1925Protoceratops andrewsi Gregory and Mook p. 1
1927Protoceratops andrewsi Huene p. 272
1928Protoceratops andrewsi Van Straelen p. 307
1929Protoceratops andrewsi Sowerby p. 149
1930Protoceratops andrewsi Osborn p. 534
1933Protoceratops andrewsi Lull p. 74
1935Protoceratops andrewsi Young
1936Protoceratops andrewsi Granger p. 22
1940Protoceratops andrewsi Brown and Schlaikjer pp. 2-3 figs. 1A, 2G-I
1942Protoceratops andrewsi Brown and Schlaikjer p. 6 fig. 2A
1946Protoceratops andrewsi Gray p. 805
1949Protoceratops andrewsi Lull and Gray p. 493
1950Protoceratops andrewsi Anonymous p. 1083
1950Protoceratops andrewsi Swinton p. 1083
1951Protoceratops andrewsi Sternberg p. 226
1953Protoceratops andrewsi Bohlin p. 13
1961Protoceratops andrewsi Schwarz et al. p. 347
1964Protoceratops andrewsi Kuhn p. 54
1964Protoceratops andrewsi Tatarinov p. 581
1965Protoceratops andrewsi Kielan-Jaworowska and Kowalski p. 178
1965Protoceratops andrewsi Rozhdestvensky
1966Protoceratops andrewsi Anonymous p. 113
1967Protoceratops andrewsi Charig p. 717
1969Protoceratops andrewsi Kielan-Jaworowska p. 33
1969Protoceratops andrewsi Schuyf p. 70
1970Protoceratops andrewsi Swinton p. 257 fig. 60
1972Protoceratops andrewsi Kurzanov p. 104
1974Protoceratops andrewsi Barsbold p. 82
1974Protoceratops andrewsi Rozhdestvensky p. 147
1977Protoceratops andrewsi Gradzinski et al. p. 301
1977Protoceratops andrewsi Rozhdestvensky p. 114
1980Protoceratops andrewsi Chao
1980Protoceratops andrewsi Coombs p. 381
1981Protoceratops andrewsi Borgomanero and Leonardi p. 217
1981Protoceratops andrewsi McIntosh p. 40
1982Protoceratops andrewsi Coombs, Jr. p. 102
1982Protoceratops andrewsi McGinnis p. 94
1984Protoceratops andrewsi Kielan-Jaworowska p. 99
1984Protoceratops andrewsi Russell p. 33
1986Proceratops andrewsi Zhao p. 72
1990Protoceratops andrewsi Dodson and Currie p. 611
1990Protoceratops andrewsi Kurzanov p. 85
1991Protoceratops andrewsi Mikhailov p. 234
1991Protoceratops andrewsi Sabath p. 153
1992Protoceratops andrewsi Dong p. 166
1994Protoceratops andrewsi Dong and Currie p. 2252
1995Protoceratops andrewsi Wang and Zhou p. 263
1996Protoceratops andrewsi Lehman p. 505
1996Protoceratops andrewsi Tereshchenko p. 232
1997Protoceratops andrewsi Barsbold p. 449
1997Protoceratops andrewsi Mikhailov p. 60
1997Protoceratops andrewsi Osmólska p. 44
1997Protoceratops andrewsi Vickaryous and Ryan p. 491
1999Protoceratops andrewsi Norell and Makovicky p. 3
2000Protoceratops andrewsi Carpenter p. 136
2000Protoceratops andrewsi Colbert p. 218
2000Protoceratops andrewsi Sereno p. 489
2000Protoceratops andrewsi Shuvalov p. 269
2001Protoceratops andrewsi Lambert et al. p. 5
2001Protoceratops andrewsi Makovicky p. 254 fig. 18.3
2001Protoceratops andrewsi Norell et al. p. 4
2003Bainoceratops efremovi Tereshchenko and Alifanov pp. 294-295 figs. 1-4
2003Protoceratops andrewsi Tereshchenko and Alifanov p. 294
2004Protoceratops andrewsi Chinnery p. 584
2004Protoceratops andrewsi You and Dodson p. 480
2006Protoceratops andrewsi Campione and Holmes p. 1014
2007Bainoceratops efremovi Tereshchenko p. 175
2007Protoceratops andrewsi Tereshchenko p. 176
2008Protoceratops andrewsi Alifanov p. 622
2008Protoceratops andrewsi Butler et al. p. 8
2008Protoceratops andrewsi Currie et al. p. 84
2008Protoceratops andrewsi Godefroit et al. p. 432
2009Protoceratops andrewsi Butler and Sullivan p. 30
2009Protoceratops andrewsi Hieronymus et al. p. 1377
2009Protoceratops andrewsi Tanoue et al. p. 434
2010Protoceratops andrewsi Alifanov and Bolotsky p. 88
2010Protoceratops andrewsi Lee et al.
2010Protoceratops andrewsi Longrich p. 692 fig. 10
2010Bainoceratops efremovi Longrich et al. p. 955
2010Protoceratops andrewsi Longrich et al. p. 955
2010Protoceratops andrewsi Osi et al. p. 466
2010Protoceratops andrewsi Zhou et al. p. 467
2011Protoceratops andrewsi Farke p. 7
2011Protoceratops andrewsi Farke et al. p. 697
2011Protoceratops andrewsi Fastovsky et al. p. 1036
2011Protoceratops andrewsi Maidment and Barrett p. 42
2012Protoceratops andrewsi Han et al. p. 1370
2012Protoceratops andrewsi Handa et al. p. 181
2013Protoceratops andrewsi Tereshchenko and Singer p. 618
2013Protoceratops andrewsi Tykoski and Fiorillo p. 495
2014Protoceratops andrewsi Hone et al. p. 7
2014Protoceratops andrewsi Tanoue and Okazaki p. 80
2014Protoceratops andrewsi Tsogtbaatar et al. p. 109
2016Protoceratops andrewsi Currie p. 84
2017Protoceratops andrewsi Chen p. 155
2018Protoceratops andrewsi Frederickson et al. p. 2
2019Protoceratops andrewsi Herne et al. p. 572
2019Protoceratops andrewsi Morschhauser et al. p. 37
2019Protoceratops andrewsi Morschhauser et al. p. 83
2019Protoceratops andrewsi Słowiak et al. p. 1
2020Protoceratops andrewsi Czepiński p. 482
2020Bainoceratops efremovi Czepiński p. 490
2020Protoceratops andrewsi Dieudonné et al.
2022Protoceratops andrewsi Son et al. p. 3

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
classReptilia
subclassEureptilia()
—Romeriida
—Diapsida()
RankNameAuthor
—Archosauromorpha(Huene 1946)
—Crocopoda
—ArchosauriformesGauthier 1986
—Eucrocopoda
—Archosauria()
informalAvemetatarsalia
—Ornithodira
—Dinosauromorpha
—Dinosauriformes
—Dinosauria()
—Ornithischia()
—Neornithischia()
—Pyrodontia
—Cerapoda
—Marginocephalia
suborderCeratopsia
infraorderNeoceratopsia
—Coronosauria
familyProtoceratopsidae
genusProtoceratops
speciesandrewsi

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Protoceratops andrewsi Granger and Gregory 1923
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Invalid names: Bainoceratops efremovi Tereshchenko and Alifanov 2003 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yesg
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: gregariouso
Environment: terrestrialsubo
Locomotion: actively mobileo
Life habit: ground dwellingsubo
Diet: herbivoresubo
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2005-08-25 05:47:29
Modified: 2005-08-25 07:47:29
Source: g = genus, subo = suborder, o = order
References: Marsh 1875, Kiessling 2004, Norman and Weishampel 1991, Peczkis 1995

Age range: base of the Late/Upper Campanian to the top of the Campanian or 83.60000 to 72.20000 Ma

Collections (17 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Santonian - Late/Upper Campanian85.7 - 72.2Mongolia (Omnogov) Protoceratops andrewsi (167657)
Campanian83.6 - 72.2Mongolia (Ömnögovi Aimag) Protoceratops sp. (130798)
Campanian83.6 - 72.2Mongolia (Omnogov) Protoceratops sp. (129749 168808 168986 168987)
Late/Upper Campanian83.6 - 72.2Mongolia (Omnogov) Protoceratops andrewsi (22685 37009 37013 37016 type locality: 46152 118259 127668 127678) Protoceratops andrewsi, Protoceratops sp. (53229) Protoceratops sp. (55256)
Middle Campanian - Late/Upper Campanian83.6 - 72.2Mongolia (Omnogov) Protoceratops andrewsi (154402)