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Emarginula
Taxonomy
Emarginula was named by Lamarck (1801) [Sepkoski's age data: Tr Ladi-u R Sepkoski's reference number: 1066]. It is extant. Its type is Emarginula conica. It is the type genus of Emarginulini.
It was assigned to Emarginulinae by Knight et al. (1960), Pacaud and Le Renard (1995) and Harzhauser and Mandic (2001); to Fissurellidae by Stewart (1927), Weisbord (1962), Ladd (1966), Fleming (1966), Szabo (1980), Ladd (1982), Beu et al. (1990), Squires and Goedert (1994) and Todd (2001); to Emarginulidae by Martin (1904), Wilson (1993), Squires and Goedert (1995) and Lozouet et al. (2001); to Archaeogastropoda by Sepkoski (2002); and to Emarginulini by Bouchet et al. (2005).
It was assigned to Emarginulinae by Knight et al. (1960), Pacaud and Le Renard (1995) and Harzhauser and Mandic (2001); to Fissurellidae by Stewart (1927), Weisbord (1962), Ladd (1966), Fleming (1966), Szabo (1980), Ladd (1982), Beu et al. (1990), Squires and Goedert (1994) and Todd (2001); to Emarginulidae by Martin (1904), Wilson (1993), Squires and Goedert (1995) and Lozouet et al. (2001); to Archaeogastropoda by Sepkoski (2002); and to Emarginulini by Bouchet et al. (2005).
Species
E. (Altomarginula), E. (Arginula), E. (Emarginula), E. (Semperia), E. (Subzeidora), E. (Tauschia), E. abnormis, E. bella, E. clathrata, E. clathrataeformis, E. clypeata, E. conformis, E. conica (type species), E. costata, E. crassa, E. dotyhillsensis, E. elegans, E. elongata, E. fissura, E. gabbi (syn. E. radiata), E. incisura, E. kitteli, E. mareana, E. marylandica, E. muensteri, E. multiradiata, E. punctura, E. radiola, E. reticulata, E. rosea, E. squammata, E. striatissima, E. tropica, E. variegata, E. washingtoniana
Species lacking formal opinion data
Synonymy list
| Year | Name and author |
|---|---|
| 1801 | Emarginula Lamarck |
| 1904 | Emarginula Martin p. 269 |
| 1927 | Emarginula Stewart p. 313 |
| 1960 | Emarginula Knight et al. p. 226 |
| 1962 | Emarginula Weisbord p. 44 |
| 1966 | Emarginula Fleming p. 36 |
| 1966 | Emarginula Ladd p. 27 |
| 1980 | Emarginula Szabo p. 66 |
| 1982 | Emarginula Ladd p. 21 |
| 1990 | Emarginula Beu et al. p. 402 |
| 1993 | Emarginula Wilson p. 54 |
| 1994 | Emarginula Squires and Goedert p. 256 |
| 1995 | Emarginula Pacaud and Le Renard p. 152 |
| 1995 | Emarginula Squires and Goedert p. 261 |
| 2001 | Emarginula Harzhauser and Mandic p. 690 |
| 2001 | Emarginula Lozouet et al. p. 17 |
| 2001 | Emarginula Todd |
| 2002 | Emarginula Sepkoski |
| 2005 | Emarginula Bouchet et al. p. 70 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. Emarginula Lamarck 1801
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Subg. †Emarginula (Altomarginula) Haber 1932
Subg. †Emarginula (Arginula) Palmer 1937
Subg. Emarginula (Emarginula) Lamarck 1801
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Emarginula (Emarginula) bicancellata Montrouzier 1860
Emarginula (Emarginula) clypeus Adams 1852
Emarginula (Emarginula) souberbiana Pilsbry 1890
†Emarginula (Emarginula) vadanaei Toni 1912
Subg. Emarginula (Semperia) Crosse 1867
Subg. †Emarginula (Tauschia) Haber 1932
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†Emarginula (Tauschia) meneghiniana Canavari 1879
†Emarginula (Tauschia) sexagenicostae Fischer et al. 2002
†Emarginula abnormis Stoppani 1858
Emarginula bella Gabb 1865
†Emarginula clathrata Deshayes 1824
†Emarginula clathrataeformis Eichwald 1853
†Emarginula clypeata Lamarck 1802
†Emarginula conformis Cossmann and Lambert 1884
Emarginula conica Lamarck 1801
†Emarginula costata Lamarck 1802
Emarginula crassa Sowerby 1813
†Emarginula dotyhillsensis Squires and Goedert 1995
†Emarginula elegans Defrance 1819
†Emarginula elongata Defrance 1819
Emarginula fissura Linnaeus 1758
†Emarginula incisura Adams 1851
†Emarginula kitteli Dreger 1892
†Emarginula mareana Weisbord 1962
†Emarginula marylandica Martin 1904
†Emarginula muensteri Pictet 1855
†Emarginula multiradiata Weisbord 1962
†Emarginula punctura Wood 1848
†Emarginula radiola Lamarck 1802
†Emarginula reticulata Sowerby 1813
Emarginula rosea Bell 1824
†Emarginula squammata Grateloup 1836
†Emarginula striatissima Traub 1979
†Emarginula tropica Weisbord 1962
†Emarginula variegata Adams 1851
†Emarginula washingtoniana Squires and Goedert 1994
Diagnosis
No diagnoses are available
Measurements
| part | N | mean | |||||
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| height | 1 | 4.00 |
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| Source: g = genus, superf = superfamily, c = class | |||||
| References: Kiessling 2004, Hendy et al. 2009 | |||||
Age range
Maximum range based only on fossils: base of the Julian to the top of the Holocene or 235.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 232.0 Ma
Minimum age of oldest fossil (stem group age): 232.0 Ma
Collections (137 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Carnian | Hungary (Bakony) | E. muensteri (203199) | |
| Carnian | Italy | E. sp. (164191) | |
| Julian | Italy | E. goldfussii (137429) E. muensteri (34692 144631 144633) | |
| Angulata | France (Lorraine) | E. liasina (53947) | |
| Sinemurian | Italy | E. meneghiniana, E. sexagenicostae (80016) | |
| Late/Upper Sinemurian | Hungary | E. lepsiusi (118984) | |
| Raricostatum - Davoei | Hungary | E. vadanaei (78690) | |
| Pliensbachian | Italy (Veneto) | E. vedanae (98398) | |
| Pliensbachian | Hungary | E. vadanaei (91406 91407) | |
| Davoei | Hungary | E. vadanaei (108956 109115) | |
| Jamesoni | Turkey | E. vedanae (158460 158462 200773) | |
| Early/Lower Bathonian - Late/Upper Bathonian | Poland | E. duofoveata (58281) | |
| Callovian | Germany | E. sp. (58203) | |
| Middle Oxfordian | Germany (Niedersachsen) | E. sp. (31435) | |
| Early/Lower Valanginian | Switzerland | E. valangiensis (2225) | |
| Valanginian | Switzerland | E. neocomiensis (2247) | |
| Valanginian | France | E. neocomiensis, E. villersensis (2222) | |
| Barremian | France | E. sp. (2195) | |
| Aptian | Switzerland | E. gilleroni (2229) | |
| Aptian | France | E. dubisiensis (2186) E. michaillensis (2178) | |
| Albian | Switzerland | E. argonensis, E. guerangeri, E. desori (2237) | |
| Albian | France | E. jaccardi (2168) | |
| Late/Upper Albian | France | E. entrevesiensis, E. breistrofferi (58386) | |
| Cenomanian | United Kingdom (England) | E. sp. (154013) | |
| Cenomanian | Switzerland | E. gresslyi (2165) | |
| Cenomanian | Germany (Nordrhein-Westfalen) | E. sp. 1, E. sp. 2 (94265) | |
| Cenomanian - Maastrichtian | United Kingdom (Sussex) | E. affinis (235797) | |
| Coniacian | Cameroon | E. sp. (177976) | |
| Late/Upper Campanian | USA (Delaware) | E. sp. (6171) | |
| Late/Upper Maastrichtian | Poland | E. costatostriata (27818 27822 28083) | |
| Danian | Denmark | E. coralliora (176994) | |
| Danian | Puerto Rico | E. sp. (87547) | |
| Danian | Greenland | Semperia sp. (22943) | |
| Danian | France | E. coralliora, E. ciplyensis (20816) | |
| Danian | Belgium | E. corneti, E. beclardi (3118) E. corneti, E. montensis, E. beclardi (22931) | |
| Paleocene | Austria (Salzburg) | E. striatissima (84335) | |
| Paleocene | Germany | E. beclardi (3115) E. corneti, E. beclardi (3114) | |
| Selandian | Denmark | E. sp. (35265) | |
| Selandian | Greenland | Semperia sp. (22944 22945) | |
| Thanetian | Australia (Victoria) | E. sp. (6267) | |
| Thanetian | Poland | E. beclardi (27019 27020) E. mariae (27018) | |
| Waipawan | New Zealand | E. sp. (51084) | |
| Lutetian | France | E. clathrata (171525) E. clypeata (184110 184821) E. costata (184107 184815) E. costata, E. radiola (171533) E. radiola (171534) E. radiola, E. elegans (184109) | |
| Lutetian | France (Cotentin) | E. elongata, E. reticulata (184164) | |
| Bartonian | USA (Texas) | E. sp. (7439 205414 205422 205480 205481) | |
| Bartonian | USA (North Carolina) | E. sp. (5366 5436) E. sp. a (91736) E. sp. b (91737) | |
| Bartonian | France (Oise) | E. costata (3494) | |
| Bartonian | USA (Alabama) | E. arata (90600) E. sp. (205438) | |
| Bartonian | USA (Washington) | E. dotyhillsensis (161924) | |
| Bartonian - Priabonian | Australia (Western Australia) | E. sp. (179824) | |
| Late/Upper Eocene | Australia (Western Australia) | E. sp. (48441) | |
| Runangan | New Zealand | E. sp. (64816) | |
| Early/Lower Oligocene | France (Essonne) | E. nystiana (9180 9181) | |
| Rupelian - Chattian | Italy | E. sp. (180394) | |
| Chattian | Austria (Upper Austria) | E. sp. (55237) | |
| Chattian | Denmark | E. punctulata (234515) E. sp. (111475) | |
| Aquitanian | France | E. squammata (74861) | |
| Early/Lower Miocene | Marshall Islands | E. sp. A (78032) | |
| Otaian | New Zealand (Auckland) | E. komitica, E. kaiparica (84915) | |
| Otaian | New Zealand | E. n. sp. (67890) | |
| Burdigalian | Austria | E. dujardini, E. reticulata (77402) | |
| Burdigalian | Germany | E. fissura (111599) | |
| Burdigalian | Italy | E. subclathrata, E. sp. 1, E. sp. 2 (178108) | |
| Middle Miocene | Poland | E. chemnitzi (70539) | |
| Serravallian | Cyprus | E. grateloupi (71874) | |
| Badenian | Poland (Holy Cross Mountains) | E. clathrataeformis (38823) | |
| Late/Upper Miocene | Australia (South Australia) | E. dennanti (63441) E. didactica, E. dilatoria (63439) | |
| Late/Upper Miocene | Philippines | E. planulata (70138) | |
| Early/Lower Pliocene | Venezuela (Sucre) | E. mareana (41545) E. multiradiata (41544) E. tropica (41552) | |
| Zanclean | Spain | E. solidula, E. fissura (80564) | |
| Zanclean | Dominican Republic | E. sp. (74964) | |
| Zanclean | Australia (Victoria) | E. transenna (166744) | |
| Zanclean | United Kingdom (England) | E. reticula (55827) | |
| Pliocene | Pakistan | E. sp. (55400) | |
| Late/Upper Pliocene | Jamaica | E. palia (58522 69090) E. palia, E. sp. A, E. sp. B (69094) E. sp. B (69093) | |
| Late/Upper Pliocene | United Kingdom (England) | E. crassa (84764) E. fissura (84819) | |
| Piacenzian | Spain | E. adriatica (80563) E. adriatica, E. rosea (80560) E. adriatica, E. solidula, E. tuberculosa, E. huzardii, E. fissura (80561) E. adriatica, E. solidula, E. tuberculosa, E. rosea, E. fissura (80562) E. adriatica, E. tuberculosa, E. rosea (80558) E. rosea (80559) | |
| Early/Lower Pleistocene | USA (Florida) | E. pilsbryi (79336) | |
| Early/Lower Pleistocene | Panama (Bocas) | E. sp. (79871) | |
| Early/Lower Pleistocene | Italy | E. adriatica, E. solidula, E. rosea, E. fissura (78312) | |
| Pleistocene | Vanuatu | E. sp. C (78042 78046) | |
| Calabrian | Greece | E. tuberculosa (190371) | |
| Middle Pleistocene | Spain | E. fissura (128324) E. huzardi (128384) | |
| Middle Pleistocene | USA (Florida) | E. pilsbryi (67069) | |
| Middle Pleistocene - Late/Upper Pleistocene | Morocco | E. elongata (127636) | |
| Late/Upper Pleistocene | Italy | E. sp. (49518) | |
| Late/Upper Pleistocene | Japan | E. galericulata, E. crassicostata, E. retecosa, E. japonica, E. maculata (88886) | |
| Late/Upper Pleistocene | Marshall Islands | E. peasei (43321) | |
| Holocene | Tuvalu | E. bicancellata (78035) | |
| Holocene | Marshall Islands | E. peasei, E. souverbiana, E. bicancellata, E. clypeus (43306) E. sp. B, E. bicancellata (43334) |