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Bernaya
Taxonomy
Bernaya was named by Jousseaume (1884) [Sepkoski's age data: J Tith T Ol-l Sepkoski's reference number: 121]. Its type is Cypraea media.
It was reranked as Cypraea (Bernayia) by Oppenheim (1906) and Vredenburg (1925); it was reranked as Cypraea (Bernaya) by Islamoglu and Taner (2002); it was synonymized subjectively with Cypraeorbis by Darragh (2011).
It was assigned to Cypraeorbini by Gliozzi and Malatesta (1983); to Cypraeidae by Cossmann (1891), Abbass (1967), Groves (1990) and Squires and Demetrion (1992); to Cypraea by Oppenheim (1906), Vredenburg (1925) and Islamoglu and Taner (2002); to Neotaenioglossa by Sepkoski (2002); to Bernayini by Groves (2004) and Groves (2011); and to Bernayinae by Schilder (1927), Perrilliat et al. (2003), Bouchet and Rosenberg (2016) and Groves and Squires (2023).
It was reranked as Cypraea (Bernayia) by Oppenheim (1906) and Vredenburg (1925); it was reranked as Cypraea (Bernaya) by Islamoglu and Taner (2002); it was synonymized subjectively with Cypraeorbis by Darragh (2011).
It was assigned to Cypraeorbini by Gliozzi and Malatesta (1983); to Cypraeidae by Cossmann (1891), Abbass (1967), Groves (1990) and Squires and Demetrion (1992); to Cypraea by Oppenheim (1906), Vredenburg (1925) and Islamoglu and Taner (2002); to Neotaenioglossa by Sepkoski (2002); to Bernayini by Groves (2004) and Groves (2011); and to Bernayinae by Schilder (1927), Perrilliat et al. (2003), Bouchet and Rosenberg (2016) and Groves and Squires (2023).
Synonymy list
| Year | Name and author |
|---|---|
| 1884 | Bernaya Jousseaume |
| 1891 | Bernayia Cossmann |
| 1906 | Cypraea (Bernayia) Oppenheim p. 301 |
| 1925 | Cypraea (Bernayia) Vredenburg p. 303 |
| 1927 | Bernaya Schilder |
| 1967 | Bernaya Abbass p. 64 |
| 1983 | Bernaya Gliozzi and Malatesta p. 103 |
| 1990 | Bernaya Groves p. 277 |
| 1992 | Bernaya Squires and Demetrion p. 31 |
| 2002 | Cypraea (Bernaya) Islamoglu and Taner p. 49 |
| 2002 | Bernaya Sepkoski |
| 2003 | Bernaya Perrilliat et al. p. 42 |
| 2004 | Bernaya Groves p. 46 |
| 2011 | Bernaya Groves p. 47 |
| 2016 | Bernaya Bouchet and Rosenberg |
| 2023 | Bernaya Groves and Squires p. 12 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Bernaya Jousseaume 1884
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Subg. †Bernaya (Bernaya) Jousseaume 1884
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†Bernaya (Bernaya) azevedoi Oliveira 1957
†Bernaya (Bernaya) baluchistanensis Noetling 1897
†Bernaya (Bernaya) beardi Groves 2004
†Bernaya (Bernaya) burlingtonensis Schilder 1932
†Bernaya (Bernaya) coromandelica Vredenburg 1920
†Bernaya (Bernaya) crawfordcatei Groves 1990
†Bernaya (Bernaya) depereti Roman and Mazeran 1920
†Bernaya (Bernaya) ellipsoides d'Archiac and Haime 1853
†Bernaya (Bernaya) gemmellaroi Stefano 1882
†Bernaya (Bernaya) jeanae Groves 2004
†Bernaya (Bernaya) obesa Deshayes 1866
Subg. Bernaya (Protocypraea) Schilder 1927
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†Bernaya (Protocypraea) argonautica Anderson 1958
†Bernaya (Protocypraea) berryessae Anderson 1958
†Bernaya (Protocypraea) cunliffei Forbes 1846
†Bernaya (Protocypraea) globuliformis Ravn 1902
†Bernaya (Protocypraea) gualalaensis Anderson 1958
†Bernaya (Protocypraea) kayei Forbes 1846
†Bernaya (Protocypraea) kohatica Cox 1930
†Bernaya (Protocypraea) louellasaulae Groves et al. 2011
†Bernaya (Protocypraea) mississippiensis Groves 1990
†Bernaya (Protocypraea) orbignana Vredenburg 1920
†Bernaya (Protocypraea) popenoei Groves 2004
†Bernaya (Protocypraea) rineyi Groves 1990
†Bernaya (Protocypraea) sayni Schilder 1941
†Bernaya (Protocypraea) veraghoorensis Stoliczka 1867
†Bernaya grovesi Squires and Demetrion 1992
†Bernaya kaylinae Groves and Squires 2023
Bernaya media Deshayes 1835
†Bernaya quarunensis Abbass 1967
†Bernaya squiresi Groves 2011
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| L. T. Groves 1990 | Shell medium to large size, anterior end somewhat carinate, dorsum smooth, spire of medium height and partially covered, aperture wide, sides rounded, anterior and posterior canals deep, fossula smooth, concave, wide. | |
| L. T. Groves 2004 | Shell medium to large in size, anterior end somewhat carinate, dorsum smooth, spire of medium height and partially covered by successive whorls, aperture wide, sides rounded, anterior and posterior canals deep, fossula smooth, concave, wide. | |
| L. T. Groves 2011 | Shell medium to large size; anterior endsomewhat carinate; dorsum smooth; spire of mediumheight and partially covered; aperture wide, sidesrounded; anterior and posterior canals deep; fossulasmooth, concave, wide. | |
| L. T. Groves and R. L. Squires 2023 | Shell medium to large in size. Oval, sides inflated, rounded. Dorsum smooth. Spire of small to medium height and partially covered by successive whorls. Aperture wide. Anterior and posterior canals deep. Anterior end smooth, concave, and broad (Jousseaume 1884: p. 88). |
Measurements
No measurements are available
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| Source: f = family, c = class | |||||
| References: Kiessling 2004, Hendy et al. 2009 | |||||
Age range
Maximum range based only on fossils: base of the Tithonian to the top of the Bartonian or 152.10000 to 38.00000 Ma
Minimum age of oldest fossil (stem group age): 145.0 Ma
Minimum age of oldest fossil (stem group age): 145.0 Ma
Collections (46 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Tithonian | Italy (Palermo) | Cypraea gemmellaroi (179803) | |
| Late/Upper Barremian | France (Barcelonne) | Pseudocassis helveticus (179817) | |
| Cenomanian - Turonian | USA (California) | Cypraea argonautica (176025) | |
| Turonian | USA (California) | B. argonautica (176024) Cypraea berryessae (176026) | |
| Turonian | USA (Oregon) | Cypraea argonautica (86814) | |
| Late/Upper Turonian | France (Provence-Alpes-Cote d'Azur) | B. depereti (87462) | |
| Coniacian | USA (California) | Cypraea kayei (44233) | |
| Coniacian | India (Tamil Nadu) | Cypraea kayei (180663) | |
| Late/Upper Santonian - Early/Lower Campanian | Canada (British Columbia) | B. beardi (176395) | |
| Campanian | India (Tamil Nadu) | Cypraea cunliffei, Erato veraghoorensis (179034) | |
| Campanian | Brazil (Pernambuco) | Cypraea azevedoi (179815 180661) | |
| Early/Lower Campanian | USA (California) | B. jeanae (176365) B. popenoei (176369) | |
| Early/Lower Campanian - Middle Campanian | USA (Mississippi) | B. n. sp. (7456) | |
| Late/Upper Campanian | USA (California) | B. crawfordcatei (176396) B. crawfordcatei, B. gualalaensis, B. rineyi (60806) B. gualalaensis (99313 176400 176402) B. rineyi (99312) | |
| Late/Upper Campanian | USA (Delaware) | B. mortoni (6146) | |
| Middle Campanian | USA (California) | B. louellasaulae, B. popenoei (176363) | |
| Middle Campanian - Late/Upper Campanian | USA (New Jersey) | Palaeocypraea burlingtonensis (180662) | |
| Late/Upper Campanian - Early/Lower Maastrichtian | USA (California) | B. gualalaensis (176397 176399 176401) Cypraea gualalaensis (176398) | |
| Maastrichtian | USA (New Jersey) | B. burlingtonensis (78124) | |
| Early/Lower Maastrichtian | Pakistan (Balochistan) | Ovula baluchistanensis (180660) | |
| Danian | Denmark | B. globuliformis (176994) Protocypraea globuliformis (137438) | |
| Paleocene | USA (California) | B. sp. (194098) | |
| Paleocene | Germany | B. sp. (3115) | |
| Selandian - Thanetian | Pakistan (North-West Frontier) | Protocypraea kohatica (52132) | |
| Ypresian | USA (Washington) | B. grovesi (176361) | |
| Ypresian | Mexico (Baja California Sur) | B. grovesi (5496) | |
| Ypresian - Lutetian | USA (California) | B. kaylinae (8014) Eocypraea castacensis (183752) | |
| Early/Lower Eocene | China (Tibet) | Ovula ellipsoides (76993) | |
| Lutetian | Mexico (Chiapas) | B. obesa, B. media (79943) | |
| Lutetian | USA (California) | B. kaylinae (8012 173475) | |
| Bartonian | France (Île-de-France) | B. obesa (179130) | |
| Bartonian | Egypt | B. quarunensis (2960) |