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Gymnuridae
Taxonomy
Gymnuridae was named by Fowler (1934). It is extant.
It was assigned to Dasyatoidea by Cappetta (1984), Cappetta (1987); to Myliobatoidea by Kumar and Loyal (1987); to Rajiformes by Underwood (2006); to Myliobatiformes by Tabuce et al. (2005), Cicimurri and Knight (2009), Underwood and Schlogl (2013); and to Myliobatoidei by Nelson et al. (2016), van der Laan (2019), Ebersole et al. (2019), Rana et al. (2021).
It was assigned to Dasyatoidea by Cappetta (1984), Cappetta (1987); to Myliobatoidea by Kumar and Loyal (1987); to Rajiformes by Underwood (2006); to Myliobatiformes by Tabuce et al. (2005), Cicimurri and Knight (2009), Underwood and Schlogl (2013); and to Myliobatoidei by Nelson et al. (2016), van der Laan (2019), Ebersole et al. (2019), Rana et al. (2021).
Synonymy list
| Year | Name and author |
|---|---|
| 1934 | Gymnuridae Fowler |
| 1984 | Gymnuridae Cappetta |
| 1987 | Gymnuridae Cappetta p. 165 |
| 1987 | Gymnuridae Kumar and Loyal p. 68 |
| 2005 | Gymnuridae Tabuce et al. p. 387 |
| 2006 | Gymnuridae Underwood p. 234 |
| 2009 | Gymnuridae Cicimurri and Knight p. 637 |
| 2013 | Gymnuridae Underwood and Schlogl p. 501 |
| 2016 | Gymnuridae Nelson et al. p. 92 |
| 2019 | Gymnuridae Ebersole et al. p. 120 |
| 2019 | Gymnuridae van der Laan p. 45 |
| 2021 | Gymnuridae Rana et al. p. 15 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Gymnuridae Fowler 1934
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G. Gymnura Van Hasselt 1823
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Gymnura altavela Linnaeus 1758
†Gymnura delpiti Cappetta 1984
†Gymnura grootaerti Herman 1984
†Gymnura hovestadti Herman 1984
†Gymnura laterialata Werner 1989
†Gymnura micura Bloch and Schneider 1801
Invalid names: Aetoplatea Vallencienes 1841 [synonym], Gymnura transita Aver'yanov and Udovichenko 1993 [nomen nudum], Pteroplatea Muller and Henle 1837 [synonym]
G. †Pachygymnura Adnet et al. 2020
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: o = order, subp = subphylum | |||||
| References: Carroll 1988, Hendy et al. 2009, Wagner 2023 | |||||
Age range
Maximum range based only on fossils: base of the Late/Upper Cenomanian to the top of the Zanclean or 100.50000 to 3.60000 Ma
Minimum age of oldest fossil (stem group age): 93.9 Ma
Minimum age of oldest fossil (stem group age): 93.9 Ma
Collections (84 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Late/Upper Cenomanian | Egypt | Gymnura laterialata (214508) | |
| Late/Upper Maastrichtian - Danian | Morocco | Gymnuridae indet. (214373) | |
| Danian | Morocco | Gymnuridae indet. (116900 214370) | |
| Selandian | Morocco | Gymnura delpiti (133217) | |
| Selandian - Thanetian | Morocco | Gymnuridae indet., Gymnura delpiti (214839 214848) | |
| Thanetian | USA (South Carolina) | Jacquhermania duponti (186974) | |
| Thanetian | Morocco | Gymnura delpiti (214833) | |
| Thanetian | Niger | Dasyatis duponti (229530) | |
| Late/Upper Paleocene - Early/Lower Eocene | India | Gymnura sp., Subathunura casieri (91316) | |
| MP 10 | France (Picardie) | Ouledia sp., Jacquhermania duponti (68440) | |
| Ypresian | India (Rajasthan) | Gymnura sp. (277682) | |
| Ypresian | USA (South Carolina) | Jacquhermania duponti (203258) | |
| Ypresian | USA (Virginia) | Jacquhermania duponti (15595) | |
| Ypresian | Senegal | Ouledia sp. (209379) | |
| Ypresian | India (Himachal Pradesh) | Subathunura casieri (31267) | |
| Ypresian | Kyrgyzstan (Osh) | Gymnura transita (28490) | |
| Ypresian | Morocco | Ouledia sigei (214849 214850 214851) | |
| Ypresian | India | Subathunura casieri (91315) | |
| Ypresian | India (Uttarakhand) | Subathunura casieri (91312) | |
| Ypresian | United Kingdom (England) | Raja duponti (8050 13299) | |
| Ypresian - Lutetian | USA (Alabama) | Jacquhermania duponti (81179) | |
| Ypresian - Lutetian | Netherlands | Gymnura grootaerti (276494 276498) | |
| Early/Lower Eocene - Middle Eocene | United Kingdom (England) | Jacqhermania duponti (3258) | |
| Lutetian | France | Gymnura sp. 1 (277214 277215 277216) Gymnura spp. (74858) Jacquhermania sp. (277212) | |
| Lutetian | Senegal | Ouledia sp. (209380) | |
| Lutetian | Belgium | Jacquhermania duponti (242331) | |
| Lutetian | USA (Alabama) | Jacquhermania duponti (13346) Jacquhermania duponti, Gymnura sp. (276540) | |
| Lutetian | Belgium (Brussels) | Raja duponti (167775) | |
| Lutetian | United Kingdom (England) | Dasyatis duponti (41888) Gymnura latidens (41992) Gymnura sp. (41989 41996) | |
| Lutetian | Morocco | Gymnuridae indet., Ouledia sp. (91339) | |
| Lutetian | Togo | Gymnura sp. (32991) | |
| Lutetian - Bartonian | Egypt | Gymnura sp., Ouledia sp. (137540) | |
| Lutetian - Bartonian | China (Xinjiang) | Jacquhermania duponti (278018 278019) | |
| Middle Eocene | Ukraine (Crimea) | Gymnura sp. (140018) | |
| Bartonian | Myanmar | Ouledia sp. (78056) | |
| Bartonian | Myanmar (Magway Division) | Ouledia sp. (83522) | |
| Bartonian | France | Gymnura sp. 2 (161753 277209 277210) | |
| Bartonian | USA (Alabama) | Jacquhermania duponti (81266 209520) | |
| Bartonian | Tunisia (Sidi Bouzid Township) | Ouledia lacuna (173640) | |
| Bartonian | India (Rajasthan) | Ouledia lacuna (277019) | |
| Priabonian | Jordan | Gymnura sp. (32899) | |
| Priabonian | Western Sahara | Gymnura sp., Ouledia sp. (96686) | |
| Priabonian | USA (Louisiana) | Gymnura sp. (32925) | |
| Priabonian | Egypt | Gymnura sp. (137551 137568) Gymnura sp., Ouledia sp. (137550 137560 137561 137564 137569) Jacquhermania attiai, Ouledia sp. (145984) Ouledia sp. (137579 137581 137582) | |
| Chattian | Germany (Mecklenburg-Vorpommern) | Gymnura sp., Gymnura altavela (80366) | |
| Chattian | USA (South Carolina) | Gymnura sp. (97550) | |
| Burdigalian | Slovakia | Gymnura sp. (161899) | |
| Burdigalian | France | Pteroplatea sp. (276527) | |
| Barstovian | USA (California) | Pteroplatea lapislutosa (85313) | |
| Langhian | Austria (Styria) | Gymnura sp. (161934) | |
| Langhian | Mexico (Baja California) | Gymnura sp. (52183) | |
| Serravallian - Tortonian | Malta (Ras ii-Karraba) | Gymnura sp. (161384) | |
| Messinian | Portugal (Alvalade) | Gymnura sp. (65955) | |
| Messinian | Portugal | Gymnura sp. (65954 91423) | |
| Zanclean | Italy | Gymnura sp. (276542) |