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Grallator variabilis

Osteichthyes - Grallatoridae

Taxonomy
Grallator variabilis was named by Lapparent and Montenat (1967). It is considered to be a form taxon. Its type specimen is ULB-04C08_D (cast), a footprint, and it is a trace fossil. Its type locality is L'Anse de la République tracksite, F6, which is in a Rhaetian estuary/bay sandstone/claystone in France.

Sister species lacking formal opinion data
G. pisanus

Synonymy list
YearName and author
1967Grallator variabilis Lapparent and Montenat p. 16 fig. 7
1968Grallator variabilis Courel et al. p. 277
1971Grallator variabilis Haubold p. 67
1972Grallator variabilis Demathieu and Haubold p. 806
1974Grallator variabilis Haubold p. 119
1990Grallator variabilis Thulborn p. 119
1992Grallator variabilis Demathieu and Sciau p. 1561
1993Grallator variabilis Demathieu p. 232
1994Grallator variabilis Vignaud et al. p. 497
2000Grallator variabilis Gand et al. p. 617
2002Grallator variabilis Demathieu et al. p. 12
2003Grallator variabilis Demathieu p. 59
2003Grallator variabilis Montenat and Bessonnat p. 42
2005Grallator variabilis Clark et al. p. 97
2005Grallator variabilis Gand et al. p. 32
2007Grallator variabilis Bécaud p. 28
2008Grallator variabilis Bessedik et al. p. 17
2009Grallator variabilis Xing et al. p. 709
2011Grallator variabilis Chabou et al.
2012Grallator variabilis Moreau et al. p. 234
2014Grallator variabilis Moreau et al. p. 365
2017Grallator variabilis Moreau et al. p. 784
2018Grallator variabilis Moreau et al. p. 37
2019Grallator variabilis Moreau et al. p. 348
2020Grallator variables Bessedik et al. p. 30
2021Grallator variabilis Leonardi and Carvalho p. 111
2024Grallator variabilis Moreau et al. pp. 349-350

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
classReptilia
RankNameAuthor
subclassEureptilia()
—Romeriida
—Diapsida()
—Archosauromorpha(Huene 1946)
—Crocopoda
—ArchosauriformesGauthier 1986
—Eucrocopoda
—Archosauria()
informalAvemetatarsalia
—Ornithodira
—Dinosauromorpha
—Dinosauriformes
—Dinosauria()
—Saurischia()
—Theropoda()
familyGrallatoridae
genusGrallator
speciesvariabilis

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Grallator variabilis Lapparent and Montenat 1967
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Diagnosis
ReferenceDiagnosis
J. Moreau et al. 2024Tridactyl, small-sized tracks (L = 8-14 cm), longer than wide (L/W = 1.6 in average). Quite long projection of the trace of digit III (L/D = 2.5 in average). Well-defined, thin, elongated and often separated impressions of digits. Traces of digits II and III are the shortest and the longest, respectively. Base of the digito-metatarsal pad of digit IV more proximal than trace of digit II. Variable divarication angle II-IV (35° in average). Round to oval phalangeal pads. Pointed marks of claws oriented outward on digits II and IV.
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yeso
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: solitaryo
Environment: terrestrialo
Locomotion: actively mobileo
Life habit: ground dwellingo
Diet: carnivoreo
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2004-12-14 12:50:08
Modified: 2004-12-14 14:50:08
Source: o = order
References: Marsh 1875, Kiessling 2004

Age range: base of the Rhaetian to the top of the Sinemurian or 205.70000 to 192.90000 Ma

Collections (6 total)


Time interval Ma Country or state Original ID and collection number
Rhaetian205.7 - 201.4France (Pays de la Loire) Carnosauria indet. (70801) Grallator variabilis (70804) Theropoda indet. (type locality: 24798)
Rhaetian - Hettangian205.7 - 199.5Algeria (Saïda) Grallator variabilis (59340)
Hettangian201.4 - 199.5France (Region Midi-Pyrenees) Grallator variabilis (31308)
Sinemurian199.5 - 192.9France (Pays de la Loire) Grallator variabilis (200112)