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Adocus beatus

Reptilia - Testudines - Adocidae

Taxonomy
Emys beatus was named by Leidy (1865). Its type specimen is ANSP 9184, a partial shell (several neurals and costals plus a left peripheral), and it is a 3D body fossil. Its type locality is Mullica Hill, which is in a Campanian/Maastrichtian terrestrial horizon in New Jersey.

It was recombined as Adocus beatus by Cope (1868), Cope (1870), Hay (1902), Hay (1908), Hay (1930), White (1972), Meylan and Gaffney (1989), Syromyatnikova et al. (2012), Syromyatnikova et al. (2013), Syromyatnikova and Danilov (2013) and Sonoda et al. (2015).

Synonyms
  • Adocus lacer was named by Hay (1908). Its type specimen is AMNH 1350, a partial shell (most of the plastron and parts of the carapace), and it is a 3D body fossil.

    It was synonymized subjectively with Adocus beatus by White (1972).
  • Adocus punctatus was named by Marsh (1890). Its type specimen is YPM, a shell (completely restored shell), and it is a 3D body fossil. Its type locality is Hornerstown area (Hornerstown Fm.) (PROXY), which is in a Maastrichtian marginal marine marl in the Hornerstown Formation of New Jersey.

    It was synonymized subjectively with Adocus beatus by White (1972).
Synonymy list
YearName and author
1865Emys beatus Leidy p. 107 figs. Figs. 1-3, Plate XVIII
1868Adocus beatus Cope p. 235
1870Adocus beatus Cope p. 233
1890Adocus punctatus Marsh pp. 178-179 figs. Pl. 7:3
1890Emys beatus Marsh p. 179
1902Adocus beatus Hay p. 444
1904Adocus punctatus Wieland
1908Adocus punctatus Hay pp. 236-237 figs. Plate 34, figs. 6, 7; plate 35, figs, i, 2; text-figs. 296-298
1908Adocus beatus Hay p. 239 figs. Plate 34, figs. 6, 7; teit-figs. 299-301
1908Adocus lacer Hay p. 241 figs. Plate 34, fig. 8; plate 36, fig. i; text-figs. 301-307
1930Adocus beatus Hay p. 89
1930Adocus lacer Hay p. 90
1930Adocus punctatus Hay p. 90
1972Adocus beatus White, Jr. p. 2
1989Adocus beatus Meylan and Gaffney
2012Adocus beatus Syromyatnikova et al.
2013Adocus beatus Syromyatnikova et al.
2015Adocus beatus Sonoda et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
RankNameAuthor
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
superfamilyTrionychoideaFitzinger 1826
familyAdocidae
genusAdocus
speciesbeatus()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Adocus beatus Leidy 1865
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Invalid names: Adocus lacer Hay 1908 [synonym], Adocus punctatus Marsh 1890 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Body part: carapaceg
Adult length: 100 to < 1000g
Environment: freshwatersubo
Locomotion: actively mobilec
Life habit: aquaticsubo
Diet: omnivoresubo
Reproduction: oviparoussubo
Created: 2005-09-01 10:38:32
Modified: 2005-09-01 12:38:32
Source: g = genus, subo = suborder, c = class, subp = subphylum
References: Estes et al. 1969, Carroll 1988, Uetz 2005, Hendy et al. 2009

Age range: base of the Late/Upper Campanian to the top of the Late/Upper Maastrichtian or 83.50000 to 66.00000 Ma

Collections (7 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Santonian - Middle Campanian85.8 - 70.6USA (Missouri) Adocus sp. (45679)
Late/Upper Campanian83.5 - 70.6USA (New Jersey) Adocus beatus (14455)
Middle Campanian83.5 - 70.6USA (South Carolina) Adocus punctatus (180790)
Middle Campanian - Late/Upper Campanian83.5 - 70.6USA (South Carolina) Adocus punctatus (79363)
Late/Upper Campanian - Early/Lower Maastrichtian83.5 - 66.0USA (New Jersey) Emys beatus (type locality: 52504)
Late/Upper Maastrichtian70.6 - 66.0USA (New Jersey) Adocus punctatus (88034)
Late/Upper Maastrichtian - Early/Lower Paleocene70.6 - 61.7USA (New Jersey) Adocus beatus (26819)