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Barbourofelinae

Mammalia - Carnivora - Nimravidae

Synonymy list
YearName and author
1970Barbourofelinae Schultz et al.
1991Barbourofelinae Bryant p. 71
2004Barbourofelidae Morlo et al. p. 45
2007Barbourofelinae Morales et al. p. 81
2008Barbourofelinae Slater and Van Valkenburgh p. 407
2010Barbourofelinae Tseng et al.
2020Barbourofelinae Wang et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
Mammaliamorpha
Mammaliaformes
RankNameAuthor
classMammalia
Theriamorpha(Rowe 1993)
Theriiformes()
Trechnotheria
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Ferae()
CarnivoramorphaWyss and Flynn 1993
CarnivoraformesFlynn et al.
orderCarnivora
suborderFeliformiaKretzoi 1945
familyNimravidaeCope 1880
subfamilyBarbourofelinaeSchultz et al. 1970
subfamilyBarbourofelinaeSchultz et al. 1970

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Barbourofelinae Schultz et al. 1970
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Tr. †Afrosmilini Morales et al. 2001
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G. †Afrosmilus Kretzoi 1929
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Afrosmilus africanus Andrews 1914
Afrosmilus hispanicus Morales et al. 2001
Afrosmilus turkanae Schmidt-Kittler 1987
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Albanosmilus jourdani Filhol 1883
G. †Barbourofelis Schultz et al. 1970
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Barbourofelis fricki Schultz et al. 1970
Barbourofelis loveorum Baskin 1981
Barbourofelis morrisi Schultz et al. 1970
Barbourofelis piveteaui Ozansoy 1965
Barbourofelis vallesiensis de Beaumont and Crusafont-Pairo 1982
Barbourofelis whitfordi Barbour and Cook 1915
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Invalid names: Ischyrosmilus osborni Merriam 1919 [synonym]
G. †Ginsburgsmilus Morales et al. 2001
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Ginsburgsmilus napakensis Morales et al. 2001
G. †Oriensmilus Wang et al. 2020
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Oriensmilus liupanensis Wang et al. 2020
G. †Prosansanosmilus Heizmann et al. 1980
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Prosansanosmilus eggeri Morlo et al. 2004
Prosansanosmilus peregrinus Heizmann et al. 1980
G. †Sansanosmilus Kretzoi 1929
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Sansanosmilus jourdani Filhol 1883
Sansanosmilus palmidens Blainville 1841
G. †Syrtosmilus Ginsburg 1978
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Syrtosmilus syrtensis Ginsburg 1978
G. †Vampyrictis Kurten 1976
Diagnosis
ReferenceDiagnosis
M. Morlo et al. 2004Barbourofelidae are distinguished by the following: loss of P1/, M2/, P/1 and M/2; plesiomorphic tooth formula 3131/3131; apomorphic species additionally lose P2/ and P/2; scimitar-like sabreteeth with crenulations on, at least, the posterior border of upper canines; upper canines markedly compressed with vertical grooves present; strong relationships between the eruption of the upper canines and the development of other sabretooth features, especially the mandibular flange (see Peigné & de Bonis, 2003); no anterior cusp on P/3, but distinct and sometimes large posterior accessory cusp on P/3-4; protoconid of M/1 relatively tall (at least in early taxa); talonid of M/1 extremely reduced, and markedly more so than the metaconid; reduction of the talonid before the metaconid on M/1; angular chin on mandible (genial flange in the apomorphic genera) and slightly (Afrosmilus turkanae) to strongly (other taxa) curved mandibular body; short horizontal proseptum in the anteromedial corner of the auditory bulla; early and complete fusion of elements making up the bulla. In addition to those features, the Barbourofelidae differ from the Nimravidae (primitive taxa) by the following: fully ossified bulla invading the mastoid; thin wall of the caudal entotympanic, not composed of three layers as in Nimravidae; petrosal not deeply recessed in the basicranium; absence of the postglenoid foramen; presence of a parastyle on P4/; protocone on P4/located further back; bulla more anteriorly located (than in the most primitive felid genera such as Proailurus and Pseudaelurus) and consequently a more anteriorly placed foramen ovale which is close to the posterior opening of the alisphenoid canal (except Barbourofelis); a shortened palate; lateral walls of the nasopharynx converging posteriorly; broad metacarpals (known from Sansanosmilus palmidens and Barbourofelis fricki only).