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Borophagus pugnator

Mammalia - Carnivora - Canidae

Taxonomy
Porthocyon pugnator was named by Cook (1922). Its type locality is Wray, which is in a Hemphillian terrestrial horizon in Colorado.

It was recombined as Aelurodon pugnator by Matthew (1924) and Matthew and Stirton (1930); it was recombined as Osteoborus pugnator by Stirton and VanderHoof (1933), Stirton (1939), VanderHoof and Gregory (1940), Richey (1979) and Munthe (1998); it was recombined as Borophagus pugnator by VanderHoof (1931), Wang et al. (1999) and Tseng and Geisler (2016).

Synonyms
  • Osteoborus galushai was named by Webb (1969). Its type specimen is F:AM 61671, 61672, a mandible, and it is a 3D body fossil. Its type locality is Mixson's Bone Bed, which is in a Hemphillian terrestrial horizon in the Alachua Formation of Florida.

    It was synonymized subjectively with Borophagus pugnator by Wang et al. (1999).
Synonymy list
YearName and author
1922Porthocyon pugnator Cook
1924Aelurodon pugnator Matthew
1930Aelurodon pugnator Matthew and Stirton
1931Borophagus pugnator VanderHoof
1933Osteoborus pugnator Stirton and VanderHoof p. 178
1939Osteoborus pugnator Stirton
1940Osteoborus pugnator VanderHoof and Gregory
1969Osteoborus galushai Webb
1979Osteoborus pugnator Richey
1998Osteoborus pugnator Munthe
1999Borophagus pugnator Wang et al.
2016Borophagus pugnator Tseng and Geisler p. 2 figs. Table 1

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Synapsida()
Therapsida()
infraorderCynodontia()
Epicynodontia
infraorderEucynodontia
RankNameAuthor
Probainognathia
Mammaliamorpha
Mammaliaformes
classMammalia
subclassTribosphenida()
infraclassEutheria()
Placentalia
Laurasiatheria
Scrotifera
Ferae()
CarnivoramorphaWyss and Flynn 1993
CarnivoraformesFlynn et al.
orderCarnivoraBowditch 1821
suborderCaniformiaKretzoi 1943
superfamilyCanoideaSimpson 1931
familyCanidaeFischer 1817
subfamilyBorophaginaeSimpson 1945
tribeBorophaginiSimpson 1945
subtribeBorophagina
genusBorophagusCope 1892
speciespugnator(Cook 1922)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
R. A. Stirton and V. L. VanderHoof 1933Rostrum short, frontals not so abruptly elevated from level of nasals as in O. cyonoides, similar to O. littoralis; P2-3 with relatively high median cusps; no anterior and posterior cusps on P2-3 except a vestigial posterior cusp on P3; P1 with one root, P2-3 with two roots; P3 relatively long, and higher than O. cyonoides; talon of M1 similar to that in O. cyonoides and horizontal ramus not so strongly deflected outwardly below p4; p1 not vestigial, but with one root; p2 with two roots; p3 with widely separated roots, relatively high median cusp and no posterior vestigial cusp.