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Epicyon haydeni

Mammalia - Carnivora - Canidae

Taxonomy
Canis (Epicyon) haydeni was named by Leidy (1858) [type of subgenus Epicyon]. It is a 3D body fossil. Its type locality is Niobrara River (coll. Hayden 1857), which is in a Miocene terrestrial horizon in the Loop Fork Formation of Nebraska.

It was recombined as Aelurodon haydeni by Scott and Osborn (1890), Matthew (1899), Matthew and Gidley (1904), Matthew and Stirton (1930), VanderHoof and Gregory (1940), McGrew (1944), Bennett (1979) and Becker (1980); it was recombined as Epicyon haydeni by Baskin (1980), Voorhies (1990), Baskin (1998), Becker and McDonald (1998), Wang et al. (1999) and Hulbert and Whitmore (2006).

Synonyms
Synonymy list
YearName and author
1858Canis (Epicyon) haydeni Leidy p. 21
1890Aelurodon haydeni Scott and Osborn p. 66
1899Aelurodon haydeni Matthew
1904Aelurodon haydeni Matthew and Gidley p. 250
1914Tephrocyon mortifer Cook
1919Aelurodon aphobus Merriam
1930Aelurodon haydeni Matthew and Stirton
1933Osteoborus ricardoensis Stirton and VanderHoof p. 178
1933Osteoborus validus Stirton and Vanderhoof
1940Aelurodon haydeni VanderHoof and Gregory
1944Aelurodon haydeni McGrew
1957Aelurodon mortifer Kitts
1964Aelurodon mortifer Kitts
1977Aelurodon mortifer Skinner et al.
1977Aelurodon validus Skinner et al.
1979Aelurodon haydeni Bennett
1979Aelurodon validus Bennett
1979Aelurodon ricardoensis Richey
1979Aelurodon validus Richey
1980Epicyon aphobus Baskin
1980Epicyon haydeni Baskin
1980Epicyon mortifer Baskin
1980Epicyon validus Baskin
1980Aelurodon haydeni Becker
1989Epicyon validus Munthe
1990Epicyon haydeni Voorhies
1992Epicyon aphobus Whistler and Burbank
1998Epicyon haydeni Baskin
1998Epicyon haydeni Becker and McDonald
1998Epicyon validus Munthe
1999Epicyon haydeni Wang et al. p. 252
2006Epicyon haydeni Hulbert and Whitmore p. 5

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
OlfactoresJefferies 1991
subphylumVertebrata
Gnathostomata()
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Synapsida()
Therapsida()
infraorderCynodontia()
Epicynodontia
RankNameAuthor
infraorderEucynodontia
Probainognathia
Mammaliamorpha
Mammaliaformes
classMammalia
subclassTribosphenida()
infraclassEutheria()
Placentalia
Laurasiatheria
Scrotifera
Ferae()
CarnivoramorphaWyss and Flynn 1993
CarnivoraformesFlynn et al.
orderCarnivoraBowditch 1821
suborderCaniformiaKretzoi 1943
superfamilyCanoideaSimpson 1931
familyCanidaeFischer 1817
subfamilyBorophaginaeSimpson 1945
tribeBorophaginiSimpson 1945
subtribeBorophagina
genusEpicyon()
specieshaydeni()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
R. A. Stirton and V. L. VanderHoof 1933 (Osteoborus ricardoensis)Largest species of Osteoborus; Pp1 present with a single root; p2 and p3 with two roots, relatively high central cusps and faint accessory cusps; p4 with posterior border truncated obliquely along a highly upturned cingulum and with a small second (cingulum) posterior accessory cusp; tooth row relatively straight as in O. pugnator, but ramus much thicker below premolar series.